The World Spider Catalog, V15 by N. I. Platnick (c) 2000 - 2014 AMNH

Family: LYCOSIDAE

Gen. Acantholycosa Dahl, 1908 [urn:lsid:amnh.org:spidergen:01583]

N.B.: Wunderlich's (1984) synonymy of this genus with Pardosa C. L. Koch, 1847 has not been accepted by subsequent workers (see Buchar & Thaler, 1993: 328).

Transferred to other genera:
A. altaica Savelyeva, 1972 -- see Pardosa.
A. nigra (C. L. Koch, 1834) -- see Pardosa.
A. nigripalpis Giltay, 1932 -- see Pardosa.
A. pyrenaea (Simon, 1876) -- see Pyrenecosa.
A. rupicola (Dufour, 1821) -- see Pyrenecosa.
A. sibirica (Kulczyn'ski, 1908) -- see Sibirocosa.
A. spinosa (Denis, 1938) -- see Pyrenecosa.
A. subsolana (Kulczyn'ski, 1907) -- see Sibirocosa.
A. triangulata Yu & Song, 1988 -- see Mongolicosa.

In synonymy:
A. altaica Savelyeva, 1972, T from Pardosa = A. lignaria (Clerck, 1757) (Marusik, Azarkina & Koponen, 2004: 119).
A. atalanta (L. Koch, 1879, T from Pardosa) = A. norvegica (Thorell, 1872) (Holm, 1973: 99, sub Pardosa).
A. beklemischevi Charitonov, 1936 = A. norvegica (Thorell, 1872) (Mikhailov, 1997c: 122).
A. fedotovi Charitonov, 1936 = A. norvegica (Thorell, 1872) (Mikhailov, 1997c: 122).
A. foveata (Odenwall, 1901, T from Pardosa) = A. norvegica (Thorell, 1872) (Mikhailov, 1997c: 122).
A. raboti (Simon, 1887) = A. norvegica (Thorell, 1872) (Mikhailov, 1997c: 122).
A. spasskyi Charitonov, 1936 = A. norvegica (Thorell, 1872) (Mikhailov, 1997c: 122).
A. strandi Kratochvíl, 1935 = A. pedestris (Simon, 1876) (Buchar & Polenec, 1974: 82).

mf aborigenica Zyuzin & Marusik, 1988....................Russia, Mongolia [urn:lsid:amnh.org:spidersp:017190]
A. sudetica Loksa, 1965: 16, f. 23 (f, misidentified).
A. a. Zyuzin & Marusik, 1988: 1083, f. 1-6 (Dmf).
Pardosa a. Platnick, 1993c: 492.
A. a. Marusik, Azarkina & Koponen, 2004: 123, f. 108-114, 125-127, 147-151 (mf).
A. a. Marusik & Omelko, 2011: 5, f. 22, 35-36 (mf).

mf altaiensis Marusik, Azarkina & Koponen, 2004....................Russia [urn:lsid:amnh.org:spidersp:038646]
A. a. Marusik, Azarkina & Koponen, 2004: 105, f. 1-9, 56 (Dmf).
A. a. Marusik & Logunov, 2011: 58, f. 6-9, 14-18 (mf).
A. a. Marusik & Kovblyuk, 2011: 185, f. 22.19 (f).

mf azarkinae Marusik & Omelko, 2011....................Russia [urn:lsid:amnh.org:spidersp:043781]
A. a. Marusik & Omelko, 2011: 2, f. 1-5, 8-12, 26-28 (Dmf).

m azheganovae (Lobanova, 1978)....................Russia [urn:lsid:amnh.org:spidersp:017386]
Alopecosa azheganovi Lobanova, 1978: 6, f. 1 (Dm).
A. a. Marusik, Azarkina & Koponen, 2004: 128, f. 53 (Tm from Alopecosa).

mf azyuzini Marusik, Hippa & Koponen, 1996....................Russia [urn:lsid:amnh.org:spidersp:017191]
A. a. Marusik, Hippa & Koponen, 1996: 22, f. 45-50 (Dmf).
A. a. Marusik, Azarkina & Koponen, 2004: 105, f. 15-18, 24-26 (mf).

mf baltoroi (Caporiacco, 1935)....................Kashmir, Nepal, China [urn:lsid:amnh.org:spidersp:017192]
Pardosa b. Caporiacco, 1935b: 233, pl. 5, f. 17 (Dmf; N.B.: baltistana in legend).
A. b. Roewer, 1955c: 152.
A. b. Buchar, 1976: 202, f. 1-3 (mf).
A. b. Chen, Song & Kim, 1998: 72, f. 13-19 (mf).
A. b. Song, Zhu & Chen, 1999: 310, f. 186A, M (mf).
A. b. Song, Zhu & Chen, 2001: 226, f. 138A-D (mf).
A. b. Marusik, Azarkina & Koponen, 2004: 112, f. 60 (m).

f dudkoromani Marusik, Azarkina & Koponen, 2004....................Russia [urn:lsid:amnh.org:spidersp:038647]
A. d. Marusik, Azarkina & Koponen, 2004: 117, f. 80-81 (Df).

mf dudkorum Marusik, Azarkina & Koponen, 2004....................Russia [urn:lsid:amnh.org:spidersp:038648]
A. d. Marusik, Azarkina & Koponen, 2004: 113, f. 59, 73-79, 84-85, 98-99 (Dmf).

m katunensis Marusik, Azarkina & Koponen, 2004....................Russia [urn:lsid:amnh.org:spidersp:038649]
A. k. Marusik, Azarkina & Koponen, 2004: 107, f. 21-23 (Dm).

m khakassica Marusik, Azarkina & Koponen, 2004....................Russia [urn:lsid:amnh.org:spidersp:038650]
A. k. Marusik, Azarkina & Koponen, 2004: 119, f. 100-103 (Dm).

f kurchumensis Marusik, Azarkina & Koponen, 2004....................Kazakhstan [urn:lsid:amnh.org:spidersp:038651]
A. k. Marusik, Azarkina & Koponen, 2004: 119, f. 82-83 (Df).

m levinae Marusik, Azarkina & Koponen, 2004....................Russia [urn:lsid:amnh.org:spidersp:038652]
A. l. Marusik, Azarkina & Koponen, 2004: 112, f. 61-65 (Dm).

mf lignaria (Clerck, 1757)....................Palearctic [urn:lsid:amnh.org:spidersp:017193]
Araneus lignarius Clerck, 1757: 90, pl. 4, f. 4 (Dmf).
Aranea l. Olivier, 1789: 217.
Lycosa l. Sundevall, 1833a: 174.
Lycosa borealis Sundevall, 1833a: 180 (Df).
Pardosa l. Simon, 1876a: 355.
Lycosa l. Chyzer & Kulczyn'ski, 1891: 59, pl. 2, f. 21 (mf).
A. l. Dahl, 1908: 367, 369, f. 61 (mf).
A. l. Charitinov, 1926b: 59, f. 2 (f).
A. l. Dahl & Dahl, 1927: 11, f. 18, 20 (mf).
A. l. Palmgren, 1939: 30, f. 30, 33, 35 (mf).
A. l. Holm, 1947: 37, pl. 8, f. 82-83, pl. 10, f. 47 (mf).
A. l. Azheganova, 1971: 12, f. 18 (mf).
A. l. Fuhn & Niculescu-Burlacu, 1971: 60, f. 24a-e (mf).
A. l. Miller, 1971: 156, pl. XXIV, f. 12-13 (mf).
Acantholycosa altaica Savelyeva, 1972b: 454, f. 1a-b (Df).
A. l. Zyuzin & Marusik, 1988: 1085, f. 7 (m).
A. l. Heimer & Nentwig, 1991: 310, f. 830 (mf).
A. l. Buchar & Thaler, 1993: 339 (Tmf from Pardosa).
Pardosa altaica Mikhailov, 1996: 105 (Tf from Acantholycosa).
A. l. Chen, Song & Kim, 1998: 72, f. 20-22 (f).
A. l. Song, Zhu & Chen, 1999: 316, f. 186B (f).
A. l. Marusik, Azarkina & Koponen, 2004: 119, f. 27-29, 54, 115-121 (mf, S).
A. l. Almquist, 2005: 184, f. 187a-h (mf).
A. l. Marusik & Omelko, 2011: 6, f. 23, 33-34, 37 (mf).
A. l. Pfliegler, Pfeiffer & Grabolle, 2012: 180, f. 1a-b, 2a-d (mf).

mf logunovi Marusik, Azarkina & Koponen, 2004....................Russia [urn:lsid:amnh.org:spidersp:038653]
A. l. Marusik, Azarkina & Koponen, 2004: 131, f. 88-91, 173-176 (Dm, not f, f. 129, 177-180, =A. norvegica).
A. l. Fomichev & Marusik, 2011a: 65, f. 1-4, 10-13 (m, Df).

mf mordkovitchi Marusik, Azarkina & Koponen, 2004....................Russia [urn:lsid:amnh.org:spidersp:038654]
A. m. Marusik, Azarkina & Koponen, 2004: 123, f. 58, 86-87, 130-134 (Dmf).

mf norvegica (Thorell, 1872)....................Palearctic [urn:lsid:amnh.org:spidersp:017194]
Lycosa n. Thorell, 1872a: 296 (Df).
Pardosa atalanta L. Koch, 1879c: 105, pl. 3, f. 18 (Df).
Pardosa raboti Simon, 1887f: 458 (Df).
Lycosa foveata Odenwall, 1901: 260, f. 7 (Df).
Lycosa n. Kulczyn'ski, 1916: 39, pl. 2, f. 62 (f).
A. beklemischevi Charitonov, 1936: 27, f. 1-4 (Dmf).
A. fedotovi Charitonov, 1936: 31, f. 6-8 (Df).
A. n. Charitonov, 1936: 34.
A. spasskyi Charitonov, 1936: 35 (Df, for specimens of Kulczyn'ski, 1916, believed misidentified).
A. n. Palmgren, 1939: 31, f. 32, 34, 36-37 (f, Dm).
A. raboti Palmgren, 1939: 32, f. 31, 38 (f).
A. n. Holm, 1947: 38, f. 4a, 15-16, pl. 8, f. 84-85 (mf).
Pardosa foveata Roewer, 1955c: 163.
A. n. Buchar, 1963: 200, f. 1D, 2J-K (f).
A. n. Buchar, 1966b: 1, f. 4A-C (mf).
A. sudetica Azheganova, 1971: 13, f. 19 (mf, misidentified).
A. n. Holm, 1973: 99 (S).
A. fedotovi Izmailova, 1989: 18, f. 1 (f).
A. n. Izmailova, 1989: 19, f. 2 (f).
A. n. Buchar & Thaler, 1993: 339 (Tmf from Pardosa).
A. n. Kropf, 1996: 102, f. 1 (f).
A. n. Mikhailov, 1997c: 122 (S).
A. n. Marusik, Azarkina & Koponen, 2004: 128, f. 92-97, 122-124, 168-172, 181-182 (mf).
A. logunovi Marusik, Azarkina & Koponen, 2004: 131, f. 129, 177-180 (f, misidentified).
A. n. Almquist, 2005: 185, f. 188a-e (mf).
A. n. Marusik & Omelko, 2011: 7, f. 24-25, 31-32, 38-39 (mf).

mf norvegica sudetica (L. Koch, 1875) *....................Europe [urn:lsid:amnh.org:spidersp:017195]
Lycosa s. L. Koch, 1875c: 12, pl. 1, f. 8 (Df).
Pardosa s. Simon, 1876a: 356.
Lycosa s. Bösenberg, 1903: 385, pl. 36, f. 566 (f).
A. s. Dahl, 1908: 368, 371, f. 63 (f, Dm).
A. s. Charitonov, 1926b: 59, f. 3 (f).
A. s. Dahl & Dahl, 1927: 14, f. 23-25 (mf).
A. s. Charitonov, 1936: 31, f. 5 (f).
A. n. s. Buchar, 1963: 200, f. 1A-C, 2A-I (mf, reduced to subspecies).
A. n. s. Fuhn & Niculescu-Burlacu, 1971: 62, f. 26a-d (mf).
A. n. s. Miller, 1971: 156, pl. XXIV, f. 14, pl. LXI, f. 18 (mf).
A. s. Heimer & Nentwig, 1991: 310, f. 832 (mf).

mf oligerae Marusik, Azarkina & Koponen, 2004....................Russia [urn:lsid:amnh.org:spidersp:038655]
A. o. Marusik, Azarkina & Koponen, 2004: 126, f. 19-20, 128, 152-161 (Dmf).
A. o. Marusik & Omelko, 2011: 3, f. 6, 13-14, 20-21, 29-30, 40 (mf).

mf paraplumalis Marusik, Azarkina & Koponen, 2004....................Russia [urn:lsid:amnh.org:spidersp:038656]
A. p. Marusik, Azarkina & Koponen, 2004: 112, f. 38-43, 49-52, 55 (Dmf).
A. p. Fomichev & Marusik, 2011a: 65, f. 7-9 (f).

mf pedestris (Simon, 1876)....................Europe [urn:lsid:amnh.org:spidersp:017196]
Lycosa longipes Thorell, 1872a: 297 (Dm, preoccupied).
Pardosa p. Simon, 1876a: 356 (mf, replacement name).
A. p. Dahl, 1908: 368, 370, f. 62 (mf).
A. p. Dahl & Dahl, 1927: 13, f. 21-22 (mf).
Lycosa (A.) strandi Kratochvíl, 1935b: 12, f. 1 (Df).
Pardosa strandi Schenkel, 1936a: 326, f. 8 (f, Dm).
A. p. Lugetti & Tongiorgi, 1965: 219, f. XVIII.1-2. XIX.1-5 (removed mf from S of A. rupicola).
A. p. Buchar & Polenec, 1974: 81, f. A-E (mf, S).
A. p. Heimer & Nentwig, 1991: 310, f. 831 (mf).
A. p. Buchar & Thaler, 1993: 329 (Tmf from Pardosa).
A. p. Marusik, Azarkina & Koponen, 2004: 146, f. 272 (m).

m petrophila Marusik, Azarkina & Koponen, 2004....................Russia [urn:lsid:amnh.org:spidersp:038657]
A. p. Marusik, Azarkina & Koponen, 2004: 119, f. 104-107 (Dm).

mf plumalis Marusik, Azarkina & Koponen, 2004....................Russia [urn:lsid:amnh.org:spidersp:038658]
A. p. Marusik, Azarkina & Koponen, 2004: 109, f. 30-37, 44-48 (Dmf).
A. p. Fomichev & Marusik, 2011a: 65, f. 5-6 (f).

m sayanensis Marusik, Azarkina & Koponen, 2004....................Russia [urn:lsid:amnh.org:spidersp:038659]
A. s. Marusik, Azarkina & Koponen, 2004: 107, f. 10-14 (Dm).
A. s. Marusik & Kovblyuk, 2011: 184, f. 22.12, 15, 21 (m).

mf solituda (Levi & Levi, 1951)....................USA, Canada [urn:lsid:amnh.org:spidersp:018930]
Pardosa s. Levi & Levi, 1951: 225, f. 11, 16 (Dmf).
Pardosa s. Dondale & Redner, 1987: 13, f. 15-16, 35-36 (mf).
Pardosa s. Dondale & Redner, 1990: 175, f. 224-227 (mf).
A. s. Kronestedt & Marusik, 2002: 63, f. 1, 3-6, 9-11, 14, 19-20, 22-24 (Tmf from Pardosa).
A. s. Vogel, 2004: 106, f. 140-141 (mf).

m spinembolus Marusik, Azarkina & Koponen, 2004....................Russia [urn:lsid:amnh.org:spidersp:038660]
A. s. Marusik, Azarkina & Koponen, 2004: 125, f. 143-146 (Dm).

mf sterneri (Marusik, 1993)....................Russia, Mongolia [urn:lsid:amnh.org:spidersp:018939]
Pardosa s. Marusik, 1993a: 77, f. 1-3 (Dm).
A. s. Kronestedt & Marusik, 2002: 67, f. 2, 7-8, 12-13, 15-18, 21, 25-27 (Tm from Pardosa, after Marusik, Logunov & Koponen, 2000: 76, Df).
A. s. Marusik, Azarkina & Koponen, 2004: 113, f. 66-72 (mf).

m sundukovi Marusik, Azarkina & Koponen, 2004....................Russia [urn:lsid:amnh.org:spidersp:038661]
A. s. Marusik, Azarkina & Koponen, 2004: 128, f. 162-167 (Dm).
A. s. Marusik & Omelko, 2011: 3, f. 7, 15-19 (m).

mf tarbagataica Marusik & Logunov, 2011....................Kazakhstan [urn:lsid:amnh.org:spidersp:043850]
A. t. Marusik & Logunov, 2011: 58, f. 1-6, 10-13 (Dmf).

mf zinchenkoi Marusik, Azarkina & Koponen, 2004....................Russia, Kazakhstan [urn:lsid:amnh.org:spidersp:038662]
A. z. Marusik, Azarkina & Koponen, 2004: 123, f. 57, 135-142 (Dmf).