The World Spider Catalog, V15 by N. I. Platnick (c) 2000 - 2014 AMNH

Family: LIOCRANIDAE

Gen. Agroeca Westring, 1861 [urn:lsid:amnh.org:spidergen:02170]

N.B.: type specimens of the type species, A. linotina sensu Westring, 1861, belong to A. proxima (O. P.-Cambridge, 1871), so that species, rather than A. brunnea (Blackwall, 1833), is the type species (Kronestedt, 2009: 28).

Transferred to other genera:
A. aureoplumata Keyserling, 1879 -- see Corinnidae incertae sedis.
A. emertoni Kaston, 1938 -- see Liocranoeca.
A. exorta Drensky, 1929 -- see Robertus under Theridiidae.
A. flavipes Saito, 1939 -- see Clubiona under Clubionidae.
A. kltina Barrion & Litsinger, 1995 -- see Medmassa under Corinnidae.
A. praticola Bösenberg & Strand, 1906 -- see Prochora under Miturgidae.
A. pulcherrima O. P.-Cambridge, 1874 -- see Castianeira under Corinnidae.

In synonymy:
A. flava (Chamberlin & Woodbury, 1929, T from Rachodrassus=Talanites) = A. trivittata (Keyserling, 1887) (Kaston, 1938b: 567).
A. gaunitzi Tullgren, 1952 = A. proxima (O. P.-Cambridge, 1871) (Kronestedt, 2009: 37).
A. kalbica (Savelyeva, 1972) = A. maculata L. Koch, 1879 (Mikhailov & Marusik, 1996: 99).
A. kiranensis (Paik, 1991, T from Drassodes) = A. coreana Namkung, 1989 (Marusik & Koponen, 2000: 63).
A. liocranoides Denis, 1956 = A. annulipes Simon, 1878 (Bosmans, 1999: 26).
A. wallacei (Bryant, 1935, T from Drassyllus) = A. pratensis Emerton, 1890 (Ubick & Roth, 1973b: 6).

f agrestis Ponomarev, 2007....................Kazakhstan [urn:lsid:amnh.org:spidersp:041387]
A. a. Ponomarev, 2007b: 94, f. 37 (Df).

mf annulipes Simon, 1878....................Spain, Corsica, Sardinia, Morocco, Algeria [urn:lsid:amnh.org:spidersp:024143]
A. a. Simon, 1878a: 304 (Df).
A. a. Simon, 1932: 942, 972, f. 1454 (f, not m).
Agraeca liocranoides Denis, 1956a: 198, f. 7 (Df).
A. a. Bosmans, 1999: 25, f. 1-4 (f, Dm, S).

f bonghwaensis Seo, 2011....................Korea [urn:lsid:amnh.org:spidersp:044310]
A. b. Seo, 2011a: 99, f. 1-4 (Df).

mf brunnea (Blackwall, 1833)....................Palearctic [urn:lsid:amnh.org:spidersp:024144]
Agelena b. Blackwall, 1833: 351 (Df).
Philoica linotina C. L. Koch, 1843: 108, f. 826 (Df).
Philoica linotina Grube, 1859: 467 (Dm).
Agelena b. Blackwall, 1861a: 159, pl. 10, f. 102 (f).
A. haglundi Thorell, 1871a: 162 (Dmf).
A. haglundi Menge, 1873: 340, pl. 60, f. 194 (mf).
A. b. Simon, 1878a: 303 (m).
A. b. O. P.-Cambridge, 1879g: 35, pl. 2, f. 7a (mf).
A. haglundi Becker, 1896: 299, pl. 17, f. 10 (mf).
A. b. Becker, 1896: 300, pl. 17, f. 11 (mf).
A. b. Chyzer & Kulczyn'ski, 1897: 244, pl. 9, f. 55, 59 (mf).
A. b. Bösenberg, 1902: 261, pl. 24, f. 378 (mf).
A. b. Lessert, 1904: 393, pl. 6, f. 40 (m).
A. b. Reimoser, 1930b: 58, f. 15 (mf).
A. b. Simon, 1932: 940, 944, 971, f. 1445-1446, 1455 (mf).
A. b. Reimoser, 1937a: 81, f. 66-68 (mf).
A. b. Tullgren, 1946: 48, f. 14A-B, pl. 8, f. 93-95 (mf).
A. b. Locket & Millidge, 1951: 147, f. 75A-B, D (mf).
A. b. Locket & Millidge, 1957: 481, f. 2A (m).
A. b. Kekenbosch, 1962: 387, f. 1 (f).
A. b. Braendegaard, 1966: 198, f. 167-168 (mf).
A. b. Tyschchenko, 1971: 134, f. 342, 345 (mf).
A. b. Miller, 1971: 106, pl. XII, f. 9-10 (mf).
A. b. Locket, Millidge & Merrett, 1974: 15, f. 8B (m).
A. b. Jocqué, 1977d: 80, f. 6-7 (m).
A. b. Roberts, 1985: 88, f. 34d (mf).
A. b. Sterghiu, 1985: 136, f. 40a-d (mf).
A. b. Grimm, 1986: 25, f. 2, 9, 22, 24-25 (mf).
A. b. Izmailova, 1989: 111, f. 97 (f).
A. b. Heimer & Nentwig, 1991: 388, f. 1012 (mf).
A. b. Roberts, 1995: 137, f. (mf).
A. b. Roberts, 1998: 146, f. (mf).
A. b. Danilov, 1999: 313, f. 1A-C (mf).
A. b. Bosselaers & Jocqué, 2002: 250, f. 3D (m).
A. b. Almquist, 2006: 336, f. 292a-h (mf).
A. b. Kamura & Hayashi, 2009: 549, f. 10-12 (mf).
A. b. Ramírez, 2014: 216, f. 144E-F (m).

mf coreana Namkung, 1989....................Russia, Korea, Japan [urn:lsid:amnh.org:spidersp:024145]
A. c. Namkung, 1989: 24, f. 1-10 (Dmf).
Drassodes kiranensis Paik, 1991f: 46, f. 1-8 (Df).
A. c. Hayashi, 1992: 136, f. 9-10 (m).
A. c. Marusik & Koponen, 2000: 63 (S).
A. c. Kim & Choi, 2001: 80, f. 1-10 (mf).
A. c. Namkung, 2002: 412, f. 33.1a-c (mf).
A. c. Namkung, 2003: 414, f. 33.1a-c (mf).
A. c. Kim, Kim & Park, 2008: 15, f. 1 (mf).
A. c. Kamura & Hayashi, 2009: 549, f. 4-6 (mf).

mf cuprea Menge, 1873....................Europe to Central Asia [urn:lsid:amnh.org:spidersp:024146]
A. c. Menge, 1873: 339, pl. 59, f. 193a (Dmf).
A. pullata Thorell, 1875a: 95 (Df).
A. pullata Thorell, 1875c: 83 (Df).
A. chrysea L. Koch, 1876b: 315 (Dmf).
A. flavo-pilosa Simon, 1887c: 143 (Df).
A. chrysea Becker, 1896: 303, pl. 17, f. 13 (mf).
A. chrysea Chyzer & Kulczyn'ski, 1897: 244, pl. 9, f. 60, 72 (mf).
A. notata O. P.-Cambridge, 1902b: 30(3) (Dmf).
A. chrysea Bösenberg, 1902: 262, pl. 24, f. 381 (mf).
A. pullata Simon, 1932: 941, 972, f. 1448, 1451 (mf).
A. pullata Reimoser, 1937a: 82, f. 71-72 (mf).
A. c. Tullgren, 1946: 51, pl. 8, f. 97-99, pl. 9, f. 100 (mf).
A. c. Locket & Millidge, 1951: 151, f. 76E, 77D (mf).
A. c. Millidge & Locket, 1955: 171, f. 2F (f).
A. c. Buchar & Zdárek, 1960: 92, f. 1D (f).
A. pullata Kekenbosch, 1962: 387, f. 4 (f).
A. pullata Fuhn & Oltean, 1969: 168, pl. I, f. 1 (m).
A. pullata Tyschchenko, 1971: 134, f. 343, 346 (mf).
A. c. Miller, 1971: 106, pl. XII, f. 13-14 (mf).
A. c. Locket, Millidge & Merrett, 1974: 15, f. 9A (f).
A. c. Roberts, 1985: 90, f. 35c (mf).
A. c. Sterghiu, 1985: 141, f. 42a-d (mf).
A. pullata Grimm, 1986: 35, f. 31, 37-38 (mf).
A. pullata Heimer & Nentwig, 1991: 390, f. 1016 (mf).
A. c. Roberts, 1995: 138, f. (mf).
A. c. Roberts, 1998: 148, f. (mf).
A. c. Almquist, 2006: 338, f. 293a-f (mf).

f debilis O. P.-Cambridge, 1885....................Yarkand [urn:lsid:amnh.org:spidersp:024147]
A. d. O. P.-Cambridge, 1885b: 26, pl. 2, f. 19 (Df).

mf dentigera Kulczyn'ski, 1913....................Europe, Russia [urn:lsid:amnh.org:spidersp:024148]
A. d. Kulczyn'ski, 1913a: 26, pl. 1, f. 10 (Df).
A. d. Braun, 1967: 179, f. 1-2 (f).
A. d. Oltean, 1973a: 105, f. 1-2 (f).
A. d. Wunderlich, 1975b: 44, f. 6-10 (f, Dm).
A. d. Jocqué, 1977d: 80, f. 1-4 (mf).
A. d. Sterghiu, 1985: 139, f. 41a-c (f).
A. d. Grimm, 1986: 28, f. 28-29 (mf).
A. d. Heimer & Nentwig, 1991: 388, f. 1013 (f).
A. d. Roberts, 1998: 147, f. (mf).
A. d. Felton, Judd & Merrett, 2004: 91, f. 1-4 (mf).
A. d. Jonsson, 2005: 49, f. 1-6 (mf).
A. d. Almquist, 2006: 339, f. 294a-d (mf).

f dubiosissima (Strand, 1908)....................Peru [urn:lsid:amnh.org:spidersp:024149]
Hilke d. Strand, 1908e: 239 (Df).
A. d. Gertsch, 1935b: 10.

f flavens O. P.-Cambridge, 1885....................Yarkand [urn:lsid:amnh.org:spidersp:024151]
A. f. O. P.-Cambridge, 1885b: 27, pl. 2, f. 20 (Df).

f gangotrae Biswas & Roy, 2008....................India [urn:lsid:amnh.org:spidersp:042442]
A. g. Biswas & Roy, 2008: 50, f. 10-12 (Df).

f guttulata Simon, 1897....................Central Asia [urn:lsid:amnh.org:spidersp:024153]
A. g. Simon, 1897c: 11 (Df).

mf inopina O. P.-Cambridge, 1886....................Europe, Algeria [urn:lsid:amnh.org:spidersp:024154]
A. i. O. P.-Cambridge, 1886: 71, pl. 4, f. 4 (Dmf).
A. proxima Kulczyn'ski, 1898: 97, pl. 2, f. 72-73 (mf, misidentified).
A. i. Simon, 1932: 942, 972, f. 1453 (mf).
A. proxima Tullgren, 1946: 52, pl. 9, f. 102-103 (m, misidentified).
A. i. Locket & Millidge, 1951: 150, f. 76C-D, 77B-C (mf).
A. proxima Tullgren, 1952: 158, f. 5 (m, misidentified).
A. i. Roberts, 1985: 90, f. 35b (mf).
A. i. Grimm, 1986: 29, f. 32, 35-36 (mf).
A. i. Heimer & Nentwig, 1991: 390, f. 1014 (mf).
A. i. Roberts, 1995: 138, f. (mf).
A. i. Roberts, 1998: 147, f. (mf).
A. i. Bosmans, 1999: 29, f. 9-12 (mf).
A. i. Kronestedt, 2009: 28, f. 1-2, 5 (m).

mf kamurai Hayashi, 1992....................China, Japan [urn:lsid:amnh.org:spidersp:024155]
A. k. Hayashi, 1992: 134, f. 5-8 (Dmf).
A. k. Kamura & Hayashi, 2009: 549, f. 7-9 (mf).
A. k. Yin et al., 2012: 1066, f. 558a-c (f).

m kastoni Chamberlin & Ivie, 1944....................USA [urn:lsid:amnh.org:spidersp:024156]
A. k. Chamberlin & Ivie, 1944: 179, f. 200 (Dm).

mf lusatica (L. Koch, 1875)....................Europe to Kazakhstan [urn:lsid:amnh.org:spidersp:024159]
Liocranum lusaticum L. Koch, 1875c: 6, pl. 1, f. 4-6 (Dmf).
A. thorelli Kulczyn'ski, in Chyzer & Kulczyn'ski, 1897: 244, pl. 9, f. 57 (Dmf).
Liocranum l. Bösenberg, 1902: 259, pl. 24, f. 375 (mf).
A. l. Bösenberg, 1902: 261, pl. 24, f. 379 (mf).
A. l. Simon, 1932: 941, 944, 971, f. 1447, 1456 (mf).
Liocranum bisaticum Dyal, 1935: 216 (Df).
A. brunnea Reimoser, 1937a: 82, f. 69-70 (mf, misidentified).
A. l. Bristowe, 1939: 25, f. 1 (f).
A. l. Tullgren, 1946: 50, pl. 8, f. 96 (mf).
A. l. Locket & Millidge, 1951: 151, f. 75C, E (mf).
A. l. Locket & Millidge, 1957: 481, f. 2B (m).
A. l. Kekenbosch, 1962: 386, f. 2 (f).
A. l. Buchar, 1970: 52, f. 3A (f).
A. l. Locket, Millidge & Merrett, 1974: 15, f. 8A (m).
A. l. Jocqué, 1977d: 80, f. 5 (m).
A. l. Roberts, 1985: 88, f. 34e (mf).
A. l. Sterghiu, 1985: 138, f. 40e-g (mf).
A. l. Grimm, 1986: 30, f. 23, 26-27 (mf).
A. l. Heimer & Nentwig, 1991: 390, f. 1017 (mf).
A. l. Roberts, 1995: 137, f. (mf).
A. l. Roberts, 1998: 146, f. (mf).
A. l. Almquist, 2006: 339, f. 295a-f (mf).

mf maculata L. Koch, 1879....................Russia, Kazakhstan [urn:lsid:amnh.org:spidersp:024160]
A. m. L. Koch, 1879c: 92, pl. 3, f. 9 (Dm).
Leptodrassus kalbicus Savelyeva, 1972a: 1240, f. 9-10 (Df).
A. m. Holm, 1973: 104, f. 109-110 (m).
A. kalbica Ovtsharenko & Savelyeva, 1992: 132 (Tf from Leptodrassus).
A. m. Mikhailov & Marusik, 1996: 99 (Sf).
A. m. Danilov, 1999: 313, f. 1D-E (f).

mf maghrebensis Bosmans, 1999....................Morocco, Algeria, Tunisia [urn:lsid:amnh.org:spidersp:037863]
A. annulipes Simon, 1932: 942, 972, f. 1449 (m only, misidentified).
A. m. Bosmans, 1999: 27, f. 5-8 (Dmf).

mf makarovae Esyunin, 2008....................Russia [urn:lsid:amnh.org:spidersp:042033]
A. m. Esyunin, in Esyunin & Kazantsev, 2008: 247, f. 1-4 (Dmf).

mf minuta Banks, 1895....................USA [urn:lsid:amnh.org:spidersp:024161]
A. m. Banks, 1895d: 80 (Df).
Apostenus acutus Emerton, 1909: 220, pl. 9, f. 7 (Dm).
A. m. Kaston, 1938b: 568, f. 6-7, 11, 16 (mf).
A. m. Kaston, 1948: 384, f. 1369-1370, 1376-1377 (mf).

mf mongolica Schenkel, 1936....................Mongolia, China, Korea [urn:lsid:amnh.org:spidersp:024162]
A. m. Schenkel, 1936b: 283, f. 96 (Df).
A. m. Song et al., 1991b: 7, f. 1.1-4 (f, Dm).
A. m. Namkung, 1992: 96, f. 1-6 (mf).
A. m. Song, Zhu & Chen, 1999: 402, f. 238A-B, E-F (mf).
A. m. Hu, 2001: 304, f. 178.1-5 (mf).
A. m. Kim & Choi, 2001: 81, f. 11-16 (mf).
A. m. Namkung, 2002: 413, f. 33.2a-c (mf).
A. m. Namkung, 2003: 415, f. 33.2a-c (mf).
A. m. Kim, Kim & Park, 2008: 16, f. 2 (mf).

mf montana Hayashi, 1986....................China, Korea, Japan [urn:lsid:amnh.org:spidersp:024163]
A. m. Hayashi, 1986a: 24, f. 1-10 (Dmf).
A. m. Song et al., 1991b: 8, f. 2.1-2 (f).
A. m. Hayashi, 1992: 134, f. 1-4 (mf).
A. m. Song, Zhu & Chen, 1999: 402, f. 238C-D (f).
A. m. Kamura & Hayashi, 2009: 549, f. 1-3 (mf).
A. m. Seo, 2011a: 99, f. 5-8 (f).

mf ornata Banks, 1892....................USA, Canada, Alaska, Russia [urn:lsid:amnh.org:spidersp:024164]
A. o. Banks, 1892a: 23, pl. 1, f. 68 (Df).
A. repens Emerton, 1894: 412, pl. 2, f. 6 (Df).
A. o. Kaston, 1938b: 564, f. 3, 8, 12, 17 (f, Dm).
Rachodrassus monroensis Kaston, 1938c: 173, pl. 8, f. 1-4 (Dm).
A. o. Chickering, 1939: 56, f. 15-16 (f).
A. o. Kaston, 1945a: 5.
A. o. Kaston, 1948: 383, f. 1361-1364, 1375 (mf).
A. o. Dondale & Redner, 1982: 121, f. 11, 226-230 (mf).
A. o. Mikhailov & Marusik, 1996: 100, f. 25-26 (f).
A. o. Paquin & Dupérré, 2003: 152, f. 1669-1671 (mf).
A. o. Marusik & Kovblyuk, 2011: 179, f. 21.3-4 (m).

mf parva Bosmans, 2011....................Greece [urn:lsid:amnh.org:spidersp:043861]
A. p. Bosmans, 2011: 19, f. 11-14 (Dmf).

mf pratensis Emerton, 1890....................USA, Canada [urn:lsid:amnh.org:spidersp:024165]
A. p. Emerton, 1890: 190, pl. 6, f. 7 (Dmf).
A. p. Emerton, 1902: 11, f. 33-36 (mf).
Drassyllus wallacei Bryant, 1935a: 77, pl. 5, f. 7-8 (Df).
A. p. Kaston, 1938b: 566, f. 4, 9, 13, 18 (mf).
A. p. Chickering, 1939: 56, f. 12-14 (mf).
A. p. Kaston, 1948: 384, f. 1365-1368 (mf).
A. p. Ubick & Roth, 1973b: 6 (S).
A. p. Dondale & Redner, 1982: 120, f. 223-225 (mf).
A. p. Paquin & Dupérré, 2003: 152, f. 1672-1674 (mf).

mf proxima (O. P.-Cambridge, 1871) *....................Europe, Russia, Turkey [urn:lsid:amnh.org:spidersp:024166]
Agelena p. O. P.-Cambridge, 1871a: 415, pl. 54, f. 13 (Dmf).
Agelena p. L. Koch, 1872c: 143, pl. 2, f. 20-22 (mf).
A. brunea Menge, 1873: 341, pl. 60, f. 195 (mf, misidentified).
A. p. Simon, 1878a: 305.
A. p. O. P.-Cambridge, 1879g: 36, pl. 2, f. 7 (mf).
A. p. Becker, 1896: 302, pl. 17, f. 12 (mf).
A. p. Bösenberg, 1902: 262, pl. 24, f. 380 (mf).
A. p. Smith, 1908: 316, pl. 25, f. 3 (m).
A. p. Simon, 1932: 942, 971, f. 1450, 1452 (mf).
A. p. Reimoser, 1937a: 83, f. 73-74 (mf).
A. p. Tullgren, 1946: 52, pl. 9, f. 101 (f; not m).
A. p. Locket & Millidge, 1951: 147, f. 70A-B, 77A (mf).
A. gaunitzi Tullgren, 1952: 158, f. 6 (Dm) [urn:lsid:amnh.org:spidersp:024152].
A. p. Kekenbosch, 1962: 386, f. 3 (f).
A. p. Braendegaard, 1966: 196, f. 165-166 (mf).
A. p. Tyschchenko, 1971: 134, f. 344 (f).
A. p. Miller, 1971: 107, pl. XII, f. 11-12 (mf).
A. p. Roberts, 1985: 90, f. 35a (mf).
A. p. Sterghiu, 1985: 143, f. 42e-h (mf).
A. p. Grimm, 1986: 32, f. 30, 33-34 (mf).
A. p. Heimer & Nentwig, 1991: 390, f. 1015 (mf).
A. p. Roberts, 1995: 137, f. (mf).
A. p. Roberts, 1998: 147, f. (mf).
A. p. Danilov, 1999: 313, f. 1F, 2A-B (mf).
A. p. Almquist, 2006: 340, f. 296a-f (mf).
A. p. Kronestedt, 2009: 37, f. 3-4, 6 (m).
A. p. Özkütük et al., 2011: 229, f. 19-23 (mf).

f spinifera Kaston, 1938....................USA [urn:lsid:amnh.org:spidersp:024167]
A. spinifer Kaston, 1938b: 566, f. 2, 20 (Df).

mf trivittata (Keyserling, 1887)....................USA [urn:lsid:amnh.org:spidersp:024168]
Hilke t. Keyserling, 1887b: 444, pl. 6, f. 17 (Df).
Herpyllus oabus Chamberlin, 1919a: 246, pl. 16, f. 6 (Dm).
A. hesperia Banks, 1921: 101, f. 4 (Dmf).
Rachodrassus flavus Chamberlin & Woodbury, 1929: 134, pl. 2, f. 2 (Dmf).
A. t. Gertsch, 1935b: 10.
A. oaba Gertsch, 1935b: 10.
A. t. Kaston, 1938b: 567, f. 5, 21 (f, Sm).