The World Spider Catalog, V15 by N. I. Platnick (c) 2000 - 2014 AMNH

Family: ATYPIDAE

Gen. Atypus Latreille, 1804 [urn:lsid:amnh.org:spidergen:00003]

N.B.: considered a senior synonym of Proatypus Miller, 1947: 52, type A. muralis Bertkau, 1890 by Kraus & Baur, 1974: 88; not a senior synonym of Sphodros Walckenaer, 1835 (Gertsch & Platnick, 1980: 15, after Kraus & Baur, 1974: 88).

Transferred to other genera:
A. abboti (Walckenaer, 1835) -- see Sphodros.
A. bicolor Lucas, 1836 -- see Sphodros.
A. milberti (Walckenaer, 1837) -- see Sphodros.
A. niger Hentz, 1842 -- see Sphodros.
A. rufipes Latreille, 1829 -- see Sphodros.

In synonymy:
A. donggukensis Kim & Kim, 1996 = A. coreanus Kim, 1985 (Namkung, Kim & Lee, 2000: 304, 338).
A. thaleri (Braun, 1963) = A. piceus (Sulzer, 1776) (Kraus & Baur, 1974: 104).

mf affinis Eichwald, 1830....................Britain to Ukraine, North Africa [urn:lsid:amnh.org:spidersp:000078]
Oletera atypus Walckenaer, 1805: 7, pl. 1, f. 8 (Df, suppressed for lack of usage, see Bonnet, 1955: 813).
Oletera atypa Walckenaer, 1806: 11, pl. 6 (f).
A. a. Eichwald, 1830: 73, pl. 2, f. 19 (Df).
A. anachoreta Ausserer, 1871a: 133 (Dm).
A. blackwalli Simon, 1873b: 110, pl. 4, f. 1-5 (Df).
A. bleodonticus Simon, 1873b: 115, pl. 4, f. 10-11 (Df).
A. coriaceus Simon, 1881e: 11 (Dm).
A. cedrorum Simon, 1889f: 380 (Df).
A. a. Chyzer & Kulczyn'ski, 1897: 280, pl. 10, f. 72 (Dm).
A. a. Bösenberg, 1902: 221, pl. 20, f. 313 (mf).
A. a. major Simon, 1909b: 11 (Dm).
A. a. Simon, 1914a: 23, f. 48-49 (mf).
A. a. Berland, 1932b: 273, f. 407 (m).
A. a. Bristowe, 1933d: 292, f. 2, 4, 6, 8 (mf).
A. a. Ehlers, 1937: 257, f. 1a-8a, 9-10 (mf).
A. a. Locket & Millidge, 1951: 46, f. 17-18 (m).
A. a. Schenkel, 1953b: 8, f. 4d (m).
A. a. Wiehle, 1953: 1, f. 1-8 (mf).
A. a. Hallander, 1958: 72, f. 1-3 (m).
A. a. Cohen, 1962: 243, f. 6.2 (f).
A. a. Braendegaard, 1966: 27, f. 13-16 (m).
A. a. Niculescu-Burlacu, 1968: 90, f. 7 (m).
A. a. Loksa, 1969: 18, f. 10C-F, 11D (mf).
A. a. Kraus & Baur, 1974: 101, f. 7-8, 11-13, 22, 24-25, 29, 37-41 (mf).
A. a. Kraus, 1978: 243, f. 12 (m).
A. a. Roberts, 1985: 46, f. 12a (m).
A. a. Wunderlich, 1986: 210, f. 200-202.
A. a. Schwendinger, 1990b: 353, f. 10, 14 (mf).
A. a. Heimer, 1990: 8, f. 21-22 (mf).
A. a. Wunderlich, 1991: 8, f. 1, 3, 5, 8, 10, 16 (mf).
A. a. Heimer & Nentwig, 1991: 36, f. 72 (mf).
A. a. Roberts, 1995: 76, f. (m).
A. a. Roberts, 1998: 78, f. (m).
A. a. Hendrickx et al., 2002: 129, f. 1-3 (m).
A. a. Almquist, 2005: 27, f. 65a-g (m).
A. a. Le Peru, 2011: 61, f. 17 (f).

mf baotianmanensis Hu, 1994....................China [urn:lsid:amnh.org:spidersp:000079]
A. b. Hu, 1994: 127, f. 1-9 (Dmf).
A. b. Song, Zhu & Chen, 1999: 35, f. 15P, T-U (mf).
A. b. Yoo & Kim, 2002: 28, f. 62 (m).
A. b. Zhu et al., 2006b: 6, f. 1-9 (mf).
A. b. Zhu & Zhang, 2011: 32, f. 11A-I (mf).

mf coreanus Kim, 1985....................Korea [urn:lsid:amnh.org:spidersp:000080]
A. c. Kim, 1985b: 2, f. 1-9 (Dmf).
A. c. Schwendinger, 1990b: 355, f. 19 (f).
A. donggukensis Kim & Kim, 1996c: 57, f. 1-5 (Df).
A. c. Kim & Kim, 1996c: 58, f. 6-11 (mf).
A. c. Kim, Ji & Lee, 1999: 122, f. 1-7 (mf).
A. donggukensis Kim, Ji & Lee, 1999: 123, f. 8-12 (f).
A. c. Namkung, Kim & Lee, 2000: 304, 338 (S).
A. c. Yoo & Kim, 2002: 25, f. 5-6 (m).
A. c. Namkung, 2002: 25, f. 1.1a-c (mf).
A. c. Kim & Cho, 2002: 56, f. 1-6 (m).
A. c. Namkung, 2003: 25, f. 1.1a-c (mf).

mf dorsualis Thorell, 1897....................Myanmar, Thailand [urn:lsid:amnh.org:spidersp:000082]
A. d. Thorell, 1897a: 186 (Df).
A. d. Schwendinger, 1989a: 94, f. 2, 5-6, 11-12, 20-23 (f, Dm).
A. d. Schwendinger, 1990b: 355, f. 8 (m).

m flexus Zhu et al., 2006....................China [urn:lsid:amnh.org:spidersp:039435]
A. f. Zhu et al., 2006b: 8, f. 10-15 (Dm).

mf formosensis Kayashima, 1943....................Taiwan [urn:lsid:amnh.org:spidersp:000083]
A. f. Kayashima, 1943a: 43, pl. 5, f. 2, 4 (Dmf; specific name attributed to Kishida but apparently never described by that author).

mf heterothecus Zhang, 1985....................China [urn:lsid:amnh.org:spidersp:000084]
A. h. Zhang, 1985: 143, pl. 3, f. 1-9, pl. 4, f. 1-8 (Dmf).
A. h. Schwendinger, 1990b: 355, f. 6, 11, 21 (mf).
A. h. Chen & Gao, 1990: 19, f. 21a-c (mf).
A. h. Chen & Zhang, 1991: 35, f. 27.1-4 (f).
A. h. Song, Zhu & Chen, 1999: 35, f. 15Q, V-W (mf).
A. h. Yin, 2001b: 2, f. 5A-P (mf).
A. heterothecua Pan, 2003: 35, fp. 1-4 (f, lapsus).
A. h. Zhu et al., 2006b: 10, f. 16-25 (mf).
A. h. Zhu & Zhang, 2011: 33, f. 12A-I (mf).
A. h. Yin et al., 2012: 124, f. 9a-r (mf).

mf javanus Thorell, 1890....................Java [urn:lsid:amnh.org:spidersp:000085]
A. j. Thorell, 1890a: 413 (Df).
A. j. Kulczyn'ski, 1908d: 540 (Dm).
A. j. Schwendinger, 1990b: 355, f. 2-5, 9, 23-25 (mf).

mf karschi Dönitz, 1887....................China, Taiwan, Japan [urn:lsid:amnh.org:spidersp:000086]
A. karschii Dönitz, 1887: 9 (D).
A. k. Bösenberg & Strand, 1906: 99, pl. 5, f. 58 (Df).
A. k. Yaginuma, 1960: 22, f. 16 (m).
A. k. Yaginuma, 1971: 22, f. 16 (m).
A. k. Gertsch & Platnick, 1980: 6, f. 8.
A. k. Yaginuma & Nishikawa, 1980: 49, f. 1 (m).
A. k. Yin, Wang & Hu, 1983: 34, f. 4A (m).
A. k. Yaginuma, 1986a: 5, f. 4.1, 5.1 (m).
A. k. Yoshikura, 1987: 148, f. 20.3B, 20.12 (mf).
A. k. Chikuni, 1989b: 20, f. 1 (m).
A. k. Schwendinger, 1990b: 358, f. 12, 17 (mf).
A. k. Feng, 1990: 28, f. 3.1-7 (mf).
A. k. Song, Zhu & Chen, 1999: 35, f. 11B, 15R, 16A-B (mf).
A. k. Zhu et al., 2006b: 13, f. 26-38, 122-123 (mf).
A. k. Ono, 2009a: 84, f. 1-5 (mf).
A. k. Yin et al., 2012: 127, f. 10a-k, 3-15 (mf).

f lannaianus Schwendinger, 1989....................Thailand [urn:lsid:amnh.org:spidersp:000087]
A. l. Schwendinger, 1989a: 92, f. 7, 19 (Df).
A. l. Schwendinger, 1990b: 358, f. 22 (f).

mf largosaccatus Zhu et al., 2006....................China [urn:lsid:amnh.org:spidersp:039436]
A. l. Zhu et al., 2006b: 17, f. 39-49 (Dmf).

f ledongensis Zhu et al., 2006....................China [urn:lsid:amnh.org:spidersp:039437]
A. l. Zhu et al., 2006b: 19, f. 50-55 (Df).

mf magnus Namkung, 1986....................Russia, Korea [urn:lsid:amnh.org:spidersp:000088]
A. m. Namkung, 1986a: 29, f. 1-7 (Dm).
A. m. Logunov, 1990: 34, f. 1 (m).
A. m. Eskov & Zonstein, 1990: 355, f. 30 (m).
A. m. Kim & Kim, 1996c: 59, f. 11-15 (m).
A. m. Kim, Ji & Lee, 1999: 124, f. 13-17 (m).
A. m. Yoo & Kim, 2002: 25, f. 7-8 (m).
A. m. Namkung, 2002: 26, f. 1.2a-b (mf).
A. m. Namkung, 2003: 26, f. 1.2a-b (mf).
A. m. Marusik & Kovblyuk, 2011: 123, f. 9.7 (m).

m medius Oliger, 1999....................Russia [urn:lsid:amnh.org:spidersp:000089]
A. m. Oliger, 1999: 201, f. 1-12 (Dm).
A. m. Marusik & Kovblyuk, 2011: 123, f. 9.4, 6, 9, 11 (m).

mf muralis Bertkau, 1890....................Central Europe to Turkmenistan [urn:lsid:amnh.org:spidersp:000090]
A. piceus Herman, 1879: 210, 361, pl. 9, f. 203-204 (mf, misidentified).
A. m. Bertkau, 1890a: 78 (Df).
A. m. Chyzer & Kulczyn'ski, 1897: 279, pl. 10, f. 73 (f, Dm).
A. m. Ehlers, 1937: 257, f. 2c-8c (mf).
Proatypus m. Miller, 1947: 52, pl. I, f. 1c, 4-5 (f).
A. m. Schenkel, 1953b: 8, f. 4c (m).
A. m. Wiehle, 1953: 8, f. 14-20 (mf).
A. m. Blazejewski, 1958: 11, f. 1-3 (m).
A. m. Cohen, 1962: 239, f. 1-5, 6.1, 12 (mf).
Proatypus m. Loksa, 1969: 17, f. 10A-B, 11A (mf).
A. m. Miller, 1971: 54, pl. II, f. 4-7 (mf).
A. m. Kraus & Baur, 1974: 106, f. 10-11, 17-18, 23, 30, 42 (mf, possible junior S of A. karschi).
A. m. Zonstein & Fet, 1985: 65, f. 1 (m).
A. m. Schwendinger, 1990b: 358, f. 16, 30, 32 (f).
A. m. Wunderlich, 1991: 8, f. 1, 2, 4, 7, 9, 12 (mf).
A. m. Heimer & Nentwig, 1991: 36, f. 73 (mf).
A. m. Le Peru, 2011: 61, f. 18 (f).
A. m. Marusik & Kovblyuk, 2011: 123, f. 9.8 (f).

mf pedicellatus Zhu et al., 2006....................China [urn:lsid:amnh.org:spidersp:039438]
A. p. Zhu et al., 2006b: 21, f. 56-67 (Dmf).

mf piceus (Sulzer, 1776) *....................Europe to Moldavia, Iran [urn:lsid:amnh.org:spidersp:000091]
Aranea picea Sulzer, 1776: 254, pl. 30, f. 2 (D).
Aranea subterranea Roemer, 1789: 66, pl. 30, f. 2 (Df).
Aranea aquatica Roemer, 1789: 66 (Df).
A. sulzeri Latreille, 1806: 85, pl. 5, f. 2 (Dj).
A. sulzeri Dufour, 1820b: 109, pl. 73, f. 6 (Dm).
A. sulzeri Hahn, 1821: 1, pl. 4 (Df).
A. sulzeri Hahn, 1832: 117, f. 88 (f, Dm).
A. sulzeri C. L. Koch, 1847: 72, f. 1547-1548 (mf).
Oletera p. Lucas, 1859b: 169.
A. sulzeri Blackwall, 1861a: 14, pl. 1, f. 1 (Dmf).
A. p. Simon, 1873b: 112, pl. 4, f. 6-9 (Dmf).
A. p. O. P.-Cambridge, 1875e: 238, pl. 8, f. 2 (mf).
A. beckii O. P.-Cambridge, 1875e: 242, pl. 8, f. 1 (Dmf).
A. p. Simon, 1892a: 199, f. 134-135, 140, 142 (m).
A. p. Becker, 1896: 319, pl. 18, f. 1 (mf).
A. beckii Chyzer & Kulczyn'ski, 1897: 280, pl. 10, f. 74 (mf).
A. p. Bösenberg, 1902: 220, pl. 20, f. 312 (mf).
A. p. Lessert, 1910b: 11, f. 20 (mf).
A. p. Simon, 1914a: 22, 24, f. 46-47 (mf).
A. p. Bristowe, 1933d: 292, f. 1, 3, 5, 7 (mf).
A. p. Ehlers, 1937: 257, f. 1b-8b (mf).
A. p. Miller, 1947: 52, pl. I, f. 1a, 2-3 (f).
A. p. Schenkel, 1953b: 8, f. 4b (m).
A. p. Wiehle, 1953: 5, f. 10-13 (mf).
A. p. Bacescu, 1955: 1127, f. 2 (m).
A. p. Cohen, 1962: 243, f. 6.3 (f).
Proatypus thaleri Braun, 1963b: 111, f. 1 (Df).
A. p. von Broen, 1965b: 409, f. 1-3 (f).
Proatypus thaleri Loksa, 1969: 17, f. 11B (f).
A. p. Loksa, 1969: 18, f. 10G, 11C (mf).
A. p. Miller, 1971: 54, pl. II, f. 3 (f).
A. p. Kraus & Baur, 1974: 104, f. 9, 11, 14-16, 20-21, 26-27, 30, 43-44 (mf, S).
A. p. Gertsch & Platnick, 1979: 5, f. 15-16.
A. p. Haupt, 1983: 276, f. 1d (m).
A. p. Schwendinger, 1989a: 91, f. 13-14 (m).
A. p. Schwendinger, 1990b: 358, f. 13, 15, 26-29, 31, 33-36 (mf).
A. p. Wunderlich, 1991: 8, f. 1, 2, 4, 6, 9, 11 (mf).
A. p. Heimer & Nentwig, 1991: 36, f. 74 (f).
A. p. Roberts, 1995: 77, f. (m).
A. p. Roberts, 1998: 79, f. (m).
A. p. Hendrickx et al., 2002: 129, f. 1-3 (m).
A. p. Henrard, 2010a: 114, f. 1, 4a-b, 6a-b (m).
A. p. Le Peru, 2011: 61, f. 19 (f).

mf quelpartensis Namkung, 2002....................Korea [urn:lsid:amnh.org:spidersp:037424]
A. q. Namkung, 2002: 27, 618, f. 1.3a-b (Dmf).
A. q. Namkung, 2003: 27, f. 1.3a-b (mf).

mf sacculatus Zhu et al., 2006....................China [urn:lsid:amnh.org:spidersp:039439]
A. s. Zhu et al., 2006b: 24, f. 68-79 (Dmf).

mf sinensis Schenkel, 1953....................China [urn:lsid:amnh.org:spidersp:000092]
A. s. Schenkel, 1953b: 6, f. 3a-l, 4a (Dmf).
A. s. Song, Zhu & Chen, 1999: 35, f. 16C (m).
A. s. Song, Zhu & Chen, 2001: 49, f. 18A-C (m).
A. s. Zhu et al., 2006b: 26, f. 80-86 (m).

mf snetsingeri Sarno, 1973....................USA [urn:lsid:amnh.org:spidersp:000093]
A. s. Sarno, 1973: 38, f. 1-9 (Dmf).
A. s. Gertsch & Platnick, 1980: 11, f. 9, 13-20 (mf).
A. s. Schwendinger, 1990b: 360, f. 18 (f).

m sternosulcus Kim et al., 2006....................Korea [urn:lsid:amnh.org:spidersp:039440]
A. s. Kim et al., 2006: 171, f. 1-7 (Dm).

mf suiningensis Zhang, 1985....................China [urn:lsid:amnh.org:spidersp:000094]
A. s. Zhang, 1985: 140, pl. 2, f. 1-9 (Dmf).
A. s. Song, Zhu & Chen, 1999: 35, f. 15S, 16D-E (mf).
A. s. Zhu et al., 2006b: 28, f. 87-98 (mf).
A. s. Yin et al., 2012: 129, f. 11a-k (mf).

mf suthepicus Schwendinger, 1989....................Thailand [urn:lsid:amnh.org:spidersp:000095]
A. s. Schwendinger, 1989a: 89, f. 1, 3-4, 8-10, 15-18 (Dmf).
A. s. Schwendinger, 1990b: 361, f. 7, 20 (f).

mf sutherlandi Chennappaiya, 1935....................India [urn:lsid:amnh.org:spidersp:000096]
A. s. Chennappaiya, in Gravely, 1935a: 70, f. 1b-c (Dmf).
A. southerlandi Roewer, 1955c: 212 (lapsus).

m suwonensis Kim et al., 2006....................Korea [urn:lsid:amnh.org:spidersp:039441]
A. s. Kim et al., 2006: 174, f. 8-14 (Dm).

mf tibetensis Zhu et al., 2006....................China [urn:lsid:amnh.org:spidersp:039442]
A. t. Zhu et al., 2006b: 31, f. 99-110 (Dmf).

mf wataribabaorum Tanikawa, 2006....................Japan [urn:lsid:amnh.org:spidersp:039443]
A. w. Tanikawa, 2006c: 25, f. 1-14 (Dmf).
A. w. Ono, 2009a: 85, f. 6-7 (m).

m wii Siliwal, Kumar & Raven, 2014....................India [urn:lsid:amnh.org:spidersp:046793]
A. w. Siliwal, Kumar & Raven, 2014: 221, f. 1A-I (Dm).

mf yajuni Zhu et al., 2006....................China [urn:lsid:amnh.org:spidersp:039444]
A. y. Zhu et al., 2006b: 34, f. 111-121 (Dmf).