The World Spider Catalog, V15 by N. I. Platnick (c) 2000 - 2014 AMNH

Family: HAHNIIDAE

Gen. Hahnia C. L. Koch, 1841 [urn:lsid:amnh.org:spidergen:01941]

N.B.: considered a senior synonym of Hahnistea Chamberlin & Ivie, 1942 and Unzickeria Lehtinen, 1967: 273, type Hahnia okefenokensis Chamberlin & Ivie, 1934 by Opell & Beatty, 1976: 420, of Hahniops Roewer, 1942 by Bosmans, 1980: 94, contra Lehtinen, 1967: 237, who regarded that genus as a junior synonym of Muizenbergia Hewitt, 1915, and of Muizenbergia Hewitt, 1915 by Bosmans, 1992: 90, after Ledoux (1991: 146) (that genus itself considered a senior synonym of Simonida Schiapelli & Gerschman, 1958a: 217, a replacement name for Scotussa Simon, 1898, preoccupied, by Lehtinen, 1967: 250; for illustrations of cheliceral stridulatory ridges of four species, see Jocqué & Bosmans, 1982; the establishment of Hahniharmia Wunderlich, 2004j: 1420 (type and only species H. picta) and Spinihahnia Wunderlich, 2004j: 1421 (type and only species H. petrobia), and transfers of species to Iberina Simon, 1881 suggested by Wunderlich, 2004j: 1421 were inadequately justified to be accepted without further documentation.

Transferred to other genera:
H. ampullaria Hickman, 1948 -- see Scotospilus.
H. astrolomae Hickman, 1948 -- see Alistra.
H. berlandi Marples, 1955 -- see Alistra.
H. ernsti Simon, 1897 -- see Neohahnia.
H. fagei M. Dahl, 1937 -- see Brommella under Dictynidae.
H. hickmani Forster, 1964 -- see Kapanga.
H. ljovuschkini (Pichka, 1965) -- see Iberina.
H. maindroni Simon, 1906 -- see Scotospilus.
H. maxima Caporiacco, 1935 -- see Neoantistea.
H. solitaria (Bryant, 1935) -- see Kapanga.
H. taprobanica Simon, 1898 -- see Alistra.
H. weyersi Simon, 1899 -- see Alistra.

In synonymy:
H. annulata Lessert, 1915 = H. tabulicola Simon, 1898 (Bosmans, 1981: 204).
H. bressica Simon, 1875 = H. helveola Simon, 1875 (Harm, 1966: 360).
H. cacuminata Bösenberg, 1902 = H. montana (Blackwall, 1841) (Harm, 1966: 354).
H. cinerea seminola Gertsch, 1934 = H. cinerea Emerton, 1890 (Opell & Beatty, 1976: 423).
H. flagellifera Zhu, Chen & Sha, 1989 = H. thorntoni Brignoli, 1982 (Zhang & Zhang, 2013: 525).
H. inornata Chamberlin & Ivie, 1942 = H. ononidum Simon, 1875 (Lehtinen, 1967: 237).
H. jacksoni Roewer, 1951 = H. ononidum Simon, 1875 (Harm, 1966: 362).
H. kaestneri Roewer, 1954 = H. schubotzi Strand, 1913 (Bosmans, 1986c: 340).
H. longipes (Chamberlin & Ivie, 1942) = H. sanjuanensis Exline, 1938 (Opell & Beatty, 1976: 427).
H. macrovulva Strand, 1913 = H. schubotzi Strand, 1913 (Lehtinen, 1967: 250, sub Muizenbergia).
H. monticola Bryant, 1941 = H. glacialis Sørensen, 1898 (Opell & Beatty, 1976: 429).
H. muscicola Simon, 1875 = H. candida Simon, 1875 (Harm, 1966: 352).
H. ohlerti Bösenberg, 1902 (removed from S of H. montana) = H. pusilla C. L. Koch, 1841 (Harm, 1966: 358).
H. rectispina Kulczyn'ski, 1926 = H. glacialis Sørensen, 1898 (Holm, 1970: 201).
H. rouleti Lessert, 1915 = H. schubotzi Strand, 1913 (Bosmans, 1986c: 340).
H. yueluensis Yin & Wang, 1983 = H. thorntoni Brignoli, 1982 (Zhang & Zhang, 2013: 526).

Homonym replaced:
H. maxima Denis, 1950 (elevated from subspecies of H. rouleti) -- see H. gigantea.

Nomen nudum:
H. nipponica Kishida (Yaginuma, in Brignoli, 1983c: 505).

mf abrahami (Hewitt, 1915)....................South Africa [urn:lsid:amnh.org:spidersp:021889]
Muizenbergia a. Hewitt, 1915c: 291, f. 1A-G (Dmf).
Muizenbergia. a. Lehtinen, 1967: 454, f. 356 (f).
H. a. Bosmans, 1992: 90, f. 12 (f).

f alini Tikader, 1964....................Nepal [urn:lsid:amnh.org:spidersp:021890]
H. a. Tikader, 1964a: 259, f. 2a-c (Df).

mf arizonica Chamberlin & Ivie, 1942....................USA, Alaska [urn:lsid:amnh.org:spidersp:021891]
H. a. Chamberlin & Ivie, 1942a: 26, pl. 5, f. 54-55 (Dmf).
H. a. Opell & Beatty, 1976: 424, f. 84-88 (mf).

f banksi Fage, 1938....................Costa Rica, Panama [urn:lsid:amnh.org:spidersp:021892]
H. b. Fage, 1938a: 375, f. 5 (Df).

f barbara Denis, 1937....................Algeria [urn:lsid:amnh.org:spidersp:021893]
H. b. Denis, 1937b: 1052, pl. 3, f. 13 (Df).

mf barbata Bosmans, 1992....................Sulawesi [urn:lsid:amnh.org:spidersp:021894]
H. b. Bosmans, 1992: 84, f. 1-6 (Dmf).

mf benoiti Bosmans & Thijs, 1980....................Kenya [urn:lsid:amnh.org:spidersp:021895]
H. vangoethemi Benoit, 1978d: 621, f. 5E (f, misidentified).
H. b. Bosmans & Thijs, 1980: 564, f. 1a-b, 2b (Dmf).

mf breviducta Bosmans & Thijs, 1980....................Kenya [urn:lsid:amnh.org:spidersp:021896]
H. b. Bosmans & Thijs, 1980: 565, f. 2f, 3a-d (Dmf).

mf caeca (Georgescu & Sarbu, 1992)....................Romania [urn:lsid:amnh.org:spidersp:021897]
Iberina c. Georgescu & Sarbu, 1992: 139, f. 1-6 (Df).
H. c. Weiss & Sarbu, 1996: 164, f. 1-4 (Tf from Iberina, Dm).

m caelebs Brignoli, 1978....................Bhutan [urn:lsid:amnh.org:spidersp:021898]
H. c. Brignoli, 1978k: 51, f. 30-32 (Dm).

m cameroonensis Bosmans, 1987....................Cameroon [urn:lsid:amnh.org:spidersp:021899]
H. c. Bosmans, 1987: 45, f. 2C-D (Dm).

mf candida Simon, 1875....................Europe, North Africa, Turkey, Israel [urn:lsid:amnh.org:spidersp:021900]
H. c. Simon, 1875a: 143 (Df).
H. muscicola Simon, 1875a: 144, pl. 6, f. 1 (Dmf).
H. c. O. P.-Cambridge, 1879g: 71 (Dm).
H. rupicola Simon, 1884h: 120 (Dm).
H. montana Bösenberg, 1902: 238, pl. 21, f. 340 (mf, misidentified).
H. muscicola Dahl, 1912b: 596.
H. c. Dahl, 1937: 108, f. 12, 23 (f).
H. muscicola Dahl, 1937: 109, f. 9, 14, 25-27 (mf).
H. c. Simon, 1937: 1028, 1046, f. 1599-1600 (mf).
H. c. Locket & Millidge, 1953: 29, f. 20E-F (mf).
H. c. Harm, 1966: 352, f. 17-22 (mf, S).
Iberina c. Lehtinen, 1967: 240 (Tmf from Hahnia, rejected).
Iberina muscicola Lehtinen, 1967: 455, f. 365, 369 (Tmf from Hahnia, rejected).
H. c. Roberts, 1985: 168, f. 74d (mf).
H. c. Heimer & Nentwig, 1991: 370, f. 959 (mf).
H. c. Roberts, 1995: 257, f. (mf).
H. c. Roberts, 1998: 273, f. (mf).
H. c. Levy, 2003: 8, f. 9-15 (mf).
H. c. Marusik et al., 2011: 163, f. 13-14, 17 (f).

f carmelita Levy, 2007....................Israel [urn:lsid:amnh.org:spidersp:041103]
H. c. Levy, 2007: 15, f. 40-41 (Df).

f cervicornata Wang & Zhang, 1986....................China [urn:lsid:amnh.org:spidersp:021901]
H. c. Wang & Zhang, 1986: 51, f. A-D (Df).
H. c. Song, Zhu & Chen, 1999: 361, f. 210A-B (f).
H. c. Yin et al., 2012: 955, f. 483a-d (f).

mf chaoyangensis Zhu & Zhu, 1983....................China [urn:lsid:amnh.org:spidersp:021902]
H. c. Zhu & Zhu, 1983c: 149, f. a-g (Dmf).
H. c. Song, Zhu & Chen, 1999: 361, f. 210C-D, 211F (mf).

mf cinerea Emerton, 1890....................North America [urn:lsid:amnh.org:spidersp:021903]
H. c. Emerton, 1890: 197, pl. 7, f. 9 (Dm).
H. c. Emerton, 1902: 106, f. 252 (m, Df).
H. c. Gertsch, 1934c: 3, f. 1-4, 19-21 (mf).
H. cinerea seminola Gertsch, 1934c: 8 (Dmf).
H. cinerea seminola Chamberlin & Ivie, 1942a: 25, pl. 5, f. 51-53 (mf).
H. c. Kaston, 1948: 292, f. 931-933 (mf).
H. c. Chickering, 1963c: 67, f. 3-6 (mf).
H. c. Opell & Beatty, 1976: 423, f. 11-12, 78-83 (mf, S).
H. c. Paquin & Dupérré, 2003: 85, f. 744-746 (mf).

f clathrata Simon, 1898....................South Africa [urn:lsid:amnh.org:spidersp:021904]
H. c. Simon, 1898f: 11 (Df).
Muizenbergia c. Lehtinen, 1967: 250 (Tf from Hahnia, rejected).
H. c. Bosmans, 1981: 206, f. 3, 11 (f).

mf corticicola Bösenberg & Strand, 1906....................Russia, China, Korea, Taiwan, Japan [urn:lsid:amnh.org:spidersp:021905]
H. c. Bösenberg & Strand, 1906: 305, pl. 6, f. 66, pl. 16, f. 474 (Dmf).
H. c. Yaginuma, 1958f: 72, f. 18-19 (f).
H. c. Yaginuma, 1960: 89, f. 79 (f).
H. c. Lee, 1966: 66, f. 24d-e (f).
H. c. Lehtinen, 1967: 454, f. 358, 362 (mf).
H. c. Yaginuma, 1971: 89, f. 79 (f).
H. c. Paik, 1978e: 363, f. 163 (f).
H. c. Paik & Namkung, 1979: 57, f. 42.1-2 (mf).
H. c. Yin & Wang, 1983: 141, f. 1-5 (mf).
H. c. Hu, 1984: 209, f. 220.1-3 (mf).
H. c. Zhu et al., 1985: 126, f. 111a-c (m).
H. c. Song, 1987: 198, f. 156 (mf).
H. c. Zhang, 1987: 139, f. 116.1-6 (mf).
H. c. Chikuni, 1989b: 105, f. 1 (mf).
H. c. Chen & Gao, 1990: 118, f. 148a-c (mf).
H. c. Chen & Zhang, 1991: 194, f. 194.1-4 (mf).
H. c. Zhao, 1993: 137, f. 65a-c (mf).
H. c. Song, Zhu & Chen, 1999: 361, f. 210E-F, 211G-H (mf).
H. c. Hu, 2001: 112, f. 30.1-4 (mf).
H. c. Song, Zhu & Chen, 2001: 276, f. 173A-D (mf).
H. c. Namkung, 2002: 372, f. 26.1a-b (mf).
H. c. Kim & Cho, 2002: 237, f. 547-552 (m).
H. c. Namkung, 2003: 374, f. 26.1a-b (mf).
H. c. Ono, 2009a: 172, f. 6-13 (mf).
H. c. Zhu & Zhang, 2011: 311, f. 224A-D (mf).
H. c. Marusik & Kovblyuk, 2011: 161, f. 18.9 (m).
H. c. Yin et al., 2012: 956, f. 484a-e (mf).

mf crozetensis Hickman, 1939....................Crozet Is. [urn:lsid:amnh.org:spidersp:021906]
H. c. Hickman, 1939b: 170, f. 18-31 (Dmf).
H. c. Berland, 1947: 62, f. 14 (f).
Muizenbergia c. Lehtinen, 1967: 250 (Tmf from Hahnia, rejected).
H. c. Ledoux, 1991: 147, f. 14A-D, 15A-C (mf).

f dewittei Bosmans, 1986....................Congo [urn:lsid:amnh.org:spidersp:021907]
H. d. Bosmans, 1986c: 348, f. 6e-f (Df).

mf difficilis Harm, 1966....................Central, Eastern Europe [urn:lsid:amnh.org:spidersp:021908]
H. d. Harm, 1966: 356, f. 29-35 (Dmf).
Iberina d. Lehtinen, 1967: 240 (Tmf from Hahnia, rejected).
H. d. Miller, 1971: 180, pl. XXX, f. 8-10 (mf).
H. d. Heimer & Nentwig, 1991: 370, f. 961 (mf).
H. d. Wisniewski, Rozwalka & Wesolowska, 2013: 55, f. 1-16 (mf).

mf eburneensis Jocqué & Bosmans, 1982....................West Africa [urn:lsid:amnh.org:spidersp:021909]
H. e. Jocqué & Bosmans, 1982: 319, f. 1-5 (Dmf).

f eidmanni (Roewer, 1942)....................Bioko [urn:lsid:amnh.org:spidersp:021910]
Hahniops e. Roewer, 1942b: 250, pl. 19, f. 6 (Df).
H. e. Roewer, 1955b: 114, f. 45 (f).
Muizenbergia e. Lehtinen, 1967: 250 (Tf from Hahniops).
H. e. Bosmans, 1980: 94, f. 1-2 (Tf from Muizenbergia).
H. e. Bosmans, 1992: 90, f. 13 (f).

mf falcata Wang, 1989....................China [urn:lsid:amnh.org:spidersp:021911]
H. f. Wang, 1989b: 285, f. 1a-e (Dmf).
H. f. Song, Zhu & Chen, 1999: 361, f. 210G-H, 211I (mf).

mf flaviceps Emerton, 1913....................USA [urn:lsid:amnh.org:spidersp:021913]
H. f. Emerton, 1913b: 257, pl. 48, f. 6 (Dmf).
H. f. Gertsch, 1934c: 11, f. 24-26 (mf).
H. f. Opell & Beatty, 1976: 430, f. 110-113 (mf).

mf gigantea Bosmans, 1986....................Central Africa [urn:lsid:amnh.org:spidersp:021914]
H. rouleti maxima Denis, 1950c: 500 (Dm, preoccupied).
H. g. Bosmans, 1986c: 344, f. 1d, f, i, 2e (replacement name; Df).

mf glacialis Sørensen, 1898....................Holarctic [urn:lsid:amnh.org:spidersp:021915]
H. g. Sørensen, 1898: 219 (Dmf).
H. rectispina Kulczyn'ski, 1926: 68, pl. 3, f. 28-29 (Dm).
H. g. Braendegaard, 1937: 4, f. 1-2 (mf).
H. g. Holm, 1939a: 73, f. 2 (f).
H. monticola Bryant, 1941: 132, f. 1 (Df).
H. g. Chamberlin & Ivie, 1947b: 17, f. 9 (m).
Neoantistea g. Lehtinen, 1967: 251, f. 375 (Tmf from Hahnia, rejected).
Neoantistea monticola Lehtinen, 1967: 251 (Tf from Hahnia, rejected).
Neoantistea rectispina Lehtinen, 1967: 251 (Tm from Hahnia, rejected).
H. g. Holm, 1970: 201 (S).
H. g. Opell & Beatty, 1976: 429, f. 106-109 (mf, S).
H. g. Marusik, Hippa & Koponen, 1996: 30, f. 85-88 (mf).
H. g. Paquin & Dupérré, 2003: 85, f. 747-749 (mf).

m harmae Brignoli, 1977....................Tunisia [urn:lsid:amnh.org:spidersp:021916]
H. h. Brignoli, 1977e: 17, f. 1-2 (Dm).

f hauseri Brignoli, 1978....................Balearic Is. [urn:lsid:amnh.org:spidersp:021917]
H. h. Brignoli, 1978e: 288, f. 16 (Df).

mf helveola Simon, 1875....................Europe, Turkey [urn:lsid:amnh.org:spidersp:021918]
H. h. Simon, 1875a: 139 (Dmf).
H. bressica Simon, 1875a: 141 (Dmf).
H. h. Bösenberg, 1902: 235, pl. 21, f. 335 (mf).
H. bressica Bösenberg, 1902: 237, pl. 21, f. 338 (f).
H. bressica Dahl, 1937: 106, f. 11-12 (f).
H. h. Hammen, 1947: 288, f. 3.
H. h. Locket & Millidge, 1953: 31, f. 21B-C, f. 22B (mf).
H. h. Casemir, 1955: 42, f. 8 (f).
H. h. Harm, 1966: 360, f. 41-45 (mf, S).
H. helvola Loksa, 1969: 123, f. 80D, 83D (mf).
H. h. Miller, 1971: 180, pl. XXX, f. 1-2 (mf).
H. h. Roberts, 1985: 168, f. 74f (mf).
H. h. Heimer & Nentwig, 1991: 370, f. 963 (mf).
H. h. Roberts, 1995: 256, f. (mf).
H. h. Roberts, 1998: 272, f. (mf).
H. h. Almquist, 2005: 274, f. 258a-e (mf).
H. h. Marusik et al., 2011: 163, f. 11-12, 18 (f).

f heterophthalma Simon, 1905....................Argentina [urn:lsid:amnh.org:spidersp:021919]
H. h. Simon, 1905e: 15 (Df).

mf himalayaensis Hu & Zhang, 1990....................China, Vietnam [urn:lsid:amnh.org:spidersp:021920]
H. h. Hu & Zhang, 1990: 165, f. 1-7 (Dmf).
H. h. Song, Zhu & Chen, 1999: 361, f. 210K-L, 211L-M (mf).
H. h. Hu, 2001: 112, f. 31.1-5 (mf).
H. h. Zhang, Li & Zheng, 2011: 12, f. 10A-C, 11A-D, 12A-I, 13A-G (mf).
H. h. Zhang, Li & Pham, 2013: 346, f. 3A-E (mf).

mf inflata Benoit, 1978....................Kenya [urn:lsid:amnh.org:spidersp:021921]
H. i. Benoit, 1978e: 619, f. 5A-D (Dmf).
H. i. Bosmans & Thijs, 1980: 562, f. 2a (f).

f innupta Brignoli, 1978....................Bhutan [urn:lsid:amnh.org:spidersp:021922]
H. i. Brignoli, 1978k: 51, f. 33 (Df).

m insulana Schenkel, 1938....................Madeira [urn:lsid:amnh.org:spidersp:021923]
H. i. Schenkel, 1938a: 32, f. 11 (Dm).

f isophthalma Mello-Leitão, 1941....................Argentina [urn:lsid:amnh.org:spidersp:021924]
H. i. Mello-Leitão, 1941d: 193, pl. 12, f. 58 (Df).

f jocquei Bosmans, 1982....................Malawi [urn:lsid:amnh.org:spidersp:021925]
H. j. Bosmans, 1982a: 177, f. 2-3 (Df).

f laodiana Song, 1990....................China [urn:lsid:amnh.org:spidersp:021926]
H. l. Song, 1990: 340, f. 1A-B (Df).
H. l. Song, Zhu & Chen, 1999: 361, f. 210M-N (f).

f laticeps Simon, 1898....................South Africa [urn:lsid:amnh.org:spidersp:021927]
H. l. Simon, 1898f: 11 (Df).
Muizenbergia tabulicola Lehtinen, 1967: 250 (S, rejected).
H. l. Bosmans, 1981: 208, f. 4, 10 (f).

f lehtineni Brignoli, 1978....................Bhutan [urn:lsid:amnh.org:spidersp:021928]
H. l. Brignoli, 1978k: 47, f. 29 (Df).

mf leopoldi Bosmans, 1982....................Cameroon [urn:lsid:amnh.org:spidersp:021929]
H. l. Bosmans, 1982b: 677, f. 1-7 (Dmf).

mf liangdangensis Tang, Yang & Kim, 1996....................China [urn:lsid:amnh.org:spidersp:021930]
H. l. Tang, Yang & Kim, 1996: 67, f. 1-9 (Dmf).
H. l. Song, Zhu & Chen, 1999: 361, f. 210O-P, 211N-O (mf).

f linderi Wunderlich, 1992....................Canary Is. [urn:lsid:amnh.org:spidersp:021931]
H. l. Wunderlich, 1992a: 429, f. 632 (Df).

m lobata Bosmans, 1981....................South Africa [urn:lsid:amnh.org:spidersp:021932]
H. l. Bosmans, 1981: 209, f. 7-8 (Dm).

f maginii Brignoli, 1977....................Italy [urn:lsid:amnh.org:spidersp:021933]
H. m. Brignoli, 1977f: 78, f. 47 (Df).
H. m. Zhang & Zhang, 2013: 522, f. 1A-F, 2A-B (f).

mf major Benoit, 1978....................Kenya [urn:lsid:amnh.org:spidersp:021934]
H. m. Benoit, 1978e: 612, f. 1A-E (Dmf).
H. m. Bosmans & Thijs, 1980: 560, f. 2d (f).

m manengoubensis Bosmans, 1987....................Cameroon [urn:lsid:amnh.org:spidersp:021935]
H. m. Bosmans, 1987: 41, f. 1A-B (Dm).

f martialis Bösenberg & Strand, 1906....................Japan [urn:lsid:amnh.org:spidersp:021936]
H. m. Bösenberg & Strand, 1906: 305, pl. 16, f. 477 (Df).
H. m. Ono, 2009a: 172, f. 16 (f).

f mauensis Bosmans, 1986....................Kenya [urn:lsid:amnh.org:spidersp:021937]
H. m. Bosmans, 1986c: 346, f. 6a-b (Df).

f melloleitaoi Schiapelli & Gerschman, 1942....................Argentina [urn:lsid:amnh.org:spidersp:021938]
H. m. Schiapelli & Gerschman, 1942: 331, f. 20-22 (Df).

mf michaelseni Simon, 1902....................Chile, Argentina, Falkland Is. [urn:lsid:amnh.org:spidersp:021939]
H. m. Simon, 1902h: 39 (Df).
H. m. Vellard, 1958: 136, f. 18-25 (f, Dm).
H. m. Lehtinen, 1967: 455, f. 376 (f).
H. m. Schiapelli & Gerschman, 1974: 89, f. 34-35 (f).

f microphthalma Snazell & Duffey, 1980....................England, Switzerland, Germany, Hungary [urn:lsid:amnh.org:spidersp:021940]
H. m. Snazell & Duffey, 1980: 50, f. 1-6 (Df).
H. m. Roberts, 1985: 168, f. 74h (f).
H. m. Heimer & Nentwig, 1991: 368, f. 957 (f).
H. m. Szita et al., 1998: 342, f. 1-2 (f).
H. m. Hänggi & Stäubli, 2013: 63, f. 3 (f).

f molossidis Brignoli, 1979....................Greece [urn:lsid:amnh.org:spidersp:021941]
H. m. Brignoli, 1979b: 195, f. 28 (Df).

mf montana (Blackwall, 1841)....................Europe, Turkey, Russia [urn:lsid:amnh.org:spidersp:021942]
Agelena m. Blackwall, 1841: 622 (Df; N.B.: primary junior homonym of A. m. C. L. Koch, 1834, but that name is now a nomen dubium).
Argus montanus Walckenaer, 1847a: 505.
Agelena m. Blackwall, 1861a: 157, pl. 10, f. 100 (f, Dm).
H. m. Thorell, 1873: 435.
H. parva Kulczyn'ski, 1882b: 30, pl. 3, f. 18 (Dm).
H. cacuminata Bösenberg, 1902: 237, pl. 21, f. 339 (Df).
H. cacuminata Dahl, 1937: 108, f. 13-14 (f).
H. m. Dahl, 1937: 113, f. 17, 32 (f).
H. m. Simon, 1937: 1026, 1046, f. 1597-1598 (mf).
H. cacuminata Hull, 1951: 95, f. 1-3 (f).
H. m. Locket & Millidge, 1953: 28, f. 20B-D (mf).
H. m. Casemir, 1955: 42, f. 5-6 (mf).
H. m. Harm, 1966: 354, f. 23-28 (mf, S).
Iberina m. Lehtinen, 1967: 240 (Tmf from Hahnia, rejected).
H. m. Loksa, 1969: 121, f. 82B-C, 83C (mf).
H. m. Miller, 1971: 180, pl. XXX, f. 13-15 (mf).
H. m. Roberts, 1985: 168, f. 74c (mf).
H. m. Bürgis, 1988: 11, f. 1-8 (mf).
H. m. Heimer & Nentwig, 1991: 370, f. 960 (mf).
H. m. Roberts, 1995: 254, f. (mf).
H. m. Roberts, 1998: 270, f. (mf).
H. monata Wunderlich, 2004j: 1421, f. 38 (m, lapsus).
H. m. Almquist, 2005: 275, f. 259a-d (mf).
H. m. Marusik & Kunt, 2009a: 78, f. 1-2, 5, 9-10 (mf).
H. m. Marusik et al., 2011: 164, f. 8-10, 15 (f).
H. m. Wisniewski, Rozwalka & Wesolowska, 2013: 55, f. 17-22 (m).

f mridulae Tikader, 1970....................India [urn:lsid:amnh.org:spidersp:021943]
H. m. Tikader, 1970: 58, f. 33a-c (Df).

mf musica Brignoli, 1978....................Bhutan [urn:lsid:amnh.org:spidersp:021944]
H. m. Brignoli, 1978k: 50, f. 25, 27-28 (Dmf).

f naguaboi (Lehtinen, 1967)....................Puerto Rico [urn:lsid:amnh.org:spidersp:021945]
H. ernesti Petrunkevitch, 1929b: 78, f. 67-68 (f, misidentified, lapsus for H. ernsti).
Unzickeria n. Lehtinen, 1967: 273 (name for f of Petrunkevitch).
H. n. Brignoli, 1983c: 506.

mf nava (Blackwall, 1841)....................Palearctic [urn:lsid:amnh.org:spidersp:021946]
Agelena n. Blackwall, 1841: 622 (Dmf).
Argus navus Walckenaer, 1847a: 506.
Agelena n. Blackwall, 1861a: 158, pl. 10, f. 101 (mf).
Agelena subfusca O. P.-Cambridge, 1861a: 431 (Dm).
H. n. Simon, 1875a: 134.
H. n. Chyzer & Kulczyn'ski, 1897: 178, pl. 7, f. 13 (mf).
H. n. Bösenberg, 1902: 236, pl. 21, f. 337 (mf).
H. n. O. P.-Cambridge, 1907: 124, pl. A, f. 4-6 (mf).
H. n. Dahl, 1937: 111, f. 8, 15, 28-29 (mf).
H. n. Simon, 1937: 1031, 1045, f. 1606-1607 (mf).
H. n. Locket & Millidge, 1953: 30, f. 20A, 21A, 22A (mf).
H. n. Harm, 1966: 364, f. 51-56 (mf).
H. n. Loksa, 1969: 122, f. 80C, 82D, 83A (mf).
H. n. Fuhn & Niculescu-Burlacu, 1970: 416, f. 1-2 (m).
H. n. Miller, 1971: 180, pl. XXX, f. 5-7 (mf).
H. n. Palmgren, 1977a: 12, f. 2.5-6 (mf).
H. n. Barrientos, 1985a: 97, f. 27-30 (mf).
H. n. Roberts, 1985: 168, f. 74e (mf).
H. n. Paik, 1985e: 24, f. 1-5 (f).
H. n. Heimer & Nentwig, 1991: 372, f. 965 (mf).
H. n. Roberts, 1995: 254, f. (mf).
H. n. Weiss & Sarbu, 1996: 170, f. 8-9 (m).
H. n. Roberts, 1998: 271, f. (mf).
H. n. Namkung, 2002: 373, f. 26.2a-b (mf).
H. n. Namkung, 2003: 375, f. 26.2a-b (mf).
H. n. Lee et al., 2004: 99, unnumbered f. (m).
H. n. Almquist, 2005: 276, f. 260a-f (mf).
H. n. Ono, 2009a: 172, f. 14-15 (mf).
H. n. Marusik et al., 2011: 164, f. 6-7, 16 (f).

m nigricans Benoit, 1978....................Kenya [urn:lsid:amnh.org:spidersp:021947]
H. n. Benoit, 1978e: 615, f. 3A-D (Dm).

mf nobilis Opell & Beatty, 1976....................Mexico [urn:lsid:amnh.org:spidersp:021948]
H. n. Opell & Beatty, 1976: 428, f. 99-102 (Dmf).

m obliquitibialis Bosmans, 1982....................Malawi [urn:lsid:amnh.org:spidersp:021949]
H. o. Bosmans, 1982a: 174, f. 1 (Dm).

mf okefinokensis Chamberlin & Ivie, 1934....................USA [urn:lsid:amnh.org:spidersp:021950]
H. o. Chamberlin & Ivie, in Gertsch, 1934c: 8, f. 22-23 (Dmf).
Unzickeria o. Lehtinen, 1967: 273 (Tmf from Hahnia).
H. o. Opell & Beatty, 1976: 428, f. 103-105 (mf).
H. o. Ledoux, 2004a: 166, f. 6C-D (m).

mf ononidum Simon, 1875....................USA, Canada, Europe, Turkey, Russia [urn:lsid:amnh.org:spidersp:021951]
H. pratensis C. L. Koch, 1841a: 64, f. 639 (f, misidentified).
H. pusilla Menge, 1869: 252, pl. 48, f. 149 (Dmf; preoccupied by C. L. Koch, 1841).
H. o. Simon, 1875a: 135 (Dmf).
H. chyzeri Becker, 1886: 25 (Dm).
H. mengei Chyzer & Kulczyn'ski, 1897: 177, pl. 7, f. 14 (Dmf).
H. mengei Bösenberg, 1902: 225, f. 333 (mf).
H. mengei Lessert, 1910b: 470 (Dmf; preoccupied by Chyzer & Kulczyn'ski, 1897).
H. helveola Jackson, 1934b: 342, f. 10-12 (f, misidentified).
H. o. Simon, 1937: 1029, 1045, f. 1603-1605 (mf).
H. mengei Dahl, 1937: 105, f. 6, 10, 19-21 (mf).
H. mengei Drensky, 1942: 58, f. 21c-d (mf).
H. inornata Chamberlin & Ivie, 1942a: 26, pl. 5, f. 56-57 (Dmf).
H. jacksoni Roewer, 1951: 438 (unnecessary replacement name).
H. mengei Casemir, 1955: 42, f. 7 (f).
H. o. Harm, 1966: 362, f. 46-50 (mf, S).
H. o. Lehtinen, 1967: 237 (S).
H. mengei Hauge, 1969: 2, f. 1-2 (mf).
H. o. Loksa, 1969: 121, f. 81A, C, 83F (mf).
H. o. Miller, 1971: 180, pl. XXX, f. 3-4 (mf).
H. o. Opell & Beatty, 1976: 426, f. 89-93 (mf).
H. o. Palmgren, 1977a: 13, f. 2.10-15 (mf).
H. o. Heimer & Nentwig, 1991: 372, f. 964 (mf).
H. o. Roberts, 1998: 272, f. (mf).
H. o. Almquist, 2005: 277, f. 261a-e (mf).
H. o. Marusik & Kunt, 2009a: 78, f. 3-4, 6-8 (mf).
H. o. Marusik et al., 2011: 164, f. 3-5, 19 (f).

mf oreophila Simon, 1898....................Sri Lanka [urn:lsid:amnh.org:spidersp:021952]
H. o. Simon, 1898f: 10 (Dmf).
H. o. Simon, 1898a: 274, f. 281.
Muizenbergia o. Lehtinen, 1967: 250 (Tmf from Hahnia).

mf ovata Song & Zheng, 1982....................China [urn:lsid:amnh.org:spidersp:021953]
H. o. Song & Zheng, 1982b: 81, f. 5-8 (Df).
H. o. Hu, 1984: 210, f. 221.1-4 (f).
H. o. Song, 1987: 200, f. 158 (f).
H. o. Zhang & Wang, 1988: 205, f. 1-3 (f, Dm).
H. o. Chen & Zhang, 1991: 195, f. 195.1-4 (f).
H. o. Song, Zhu & Chen, 1999: 361, f. 210S-T, 211P-Q (mf).
H. o. Zhu & Zhang, 2011: 312, f. 225A-B (f).
H. o. Yin et al., 2012: 957, f. 485a-f (mf).

mf petrobia Simon, 1875....................Europe [urn:lsid:amnh.org:spidersp:021954]
H. p. Simon, 1875a: 132 (Dmf).
H. p. Simon, 1937: 1029, 1045, f. 1601-1602 (mf).
H. ononidum Harm, 1966: 362 (S, rejected by Barrientos).
H. p. Brignoli, 1978e: 288, f. 15 (f).
H. p. Barrientos, 1985a: 97, f. 23-26 (mf).
Spinihahnia p. Wunderlich, 2004j: 1422, f. 48-57 (mf, T not yet accepted).

mf picta Kulczyn'ski, 1897....................Europe [urn:lsid:amnh.org:spidersp:021955]
H. p. Kulczyn'ski, in Chyzer & Kulczyn'ski, 1897: 178, pl. 7, f. 15 (Df).
H. p. Kulczyn'ski, 1898: 103, pl. 2, f. 76-77 (Dm).
H. p. Simon, 1937: 1026, 1047, f. 1595-1596 (mf).
H. p. Harm, 1966: 366, f. 57-63 (mf).
H. p. Loksa, 1969: 120, f. 81B, 83A (mf).
H. p. Miller, 1971: 179, pl. XXIX, f. 25-27 (mf).
H. p. Dziabaszewski, 1975: 101, f. 2 (f).
H. p. Heimer & Nentwig, 1991: 370, f. 954 (mf).
Hahniharmia p. Wunderlich, 2004j: 1420, f. 41-47 (mf, T not yet accepted).
H. p. Pfliegler, 2014: 145, f. 1g-h, 3d, 4a (mf).

mf pinicola Arita, 1978....................Japan [urn:lsid:amnh.org:spidersp:021956]
H. p. Arita, 1978: 240, f. 1-5 (Dmf).
H. p. Ono, 2009a: 173, f. 17-19 (mf).

mf pusilla C. L. Koch, 1841 *....................Europe, Russia [urn:lsid:amnh.org:spidersp:021957]
H. p. C. L. Koch, 1841a: 61, f. 637-638 (not 639) (Dmf).
Agelena p. Canestrini & Pavesi, 1870: 42.
H. helveola Simon, 1875a: 139 (mf, in part).
H. p. Müller & Schenkel, 1895: 751, pl. 14, f. 5 (mf).
H. ohlerti Bösenberg, 1902: 234, pl. 21, f. 331 (Df).
H. p. Bösenberg, 1902: 206, pl. 21, f. 336 (mf).
H. p. O. P.-Cambridge, 1907: 124, pl. A, f. 7-11 (mf).
H. p. Jackson, 1907: 2, pl. 1, f. 1-4.
H. p. Dahl, 1937: 112, f. 7, 16, 30-31 (mf).
H. p. Simon, 1937: 1032, 1046, f. 1610-1611 (mf).
H. helveola Simon, 1937: 1032, 1046, f. 1608-1609 (mf).
H. p. Drensky, 1942: 58, f. 21a-b (mf).
H. p. Locket & Millidge, 1953: 31, f. 21D, 22C (mf).
H. p. Casemir, 1955: 42, f. 9 (f).
H. p. Harm, 1966: 358, f. 36-40 (mf, S).
H. p. Lehtinen, 1967: 454, f. 357, 359, 361 (mf).
H. p. Loksa, 1969: 124, f. 81D, 82A, 83B (mf).
H. p. Miller, 1971: 180, pl. XXX, f. 11-12 (mf).
H. p. Utochkin & Pakhorukov, 1976: 84, f. 12 (f).
H. p. Palmgren, 1977a: 13, f. 2.7-9 (mf).
H. p. Müller, 1982a: 249, f. 6 (m).
H. p. Müller, 1983a: 296, f. 1-2 (f).
H. p. Roberts, 1985: 168, f. 74g (mf).
H. p. Heimer & Nentwig, 1991: 370, f. 962 (mf).
H. p. Roberts, 1995: 256, f. (mf).
H. p. Roberts, 1998: 272, f. (mf).
H. p. Almquist, 2005: 278, f. 262a-f (mf).

mf pusio Simon, 1898....................Sri Lanka [urn:lsid:amnh.org:spidersp:021958]
H. p. Simon, 1898f: 10 (Dmf).

f pyriformis Yin & Wang, 1984....................China [urn:lsid:amnh.org:spidersp:021959]
H. p. Yin & Wang, 1984c: 269, f. 1A-C (Df).
H. p. Song, 1987: 200, f. 159 (f).
H. p. Song, Zhu & Chen, 1999: 361, f. 13G, 210U-V (f).
H. p. Yin et al., 2012: 959, f. 486a-e (f).

mf reniformis Chen, Yan & Yin, 2009....................China [urn:lsid:amnh.org:spidersp:042845]
H. r. Chen, Yan & Yin, 2009: 68, f. 7-12 (Dmf).

mf rimaformis Zhang, Li & Pham, 2013....................Vietnam [urn:lsid:amnh.org:spidersp:045723]
H. r. Zhang, Li & Pham, 2013: 347, f. 4A-F, 5A-G, 6A-G (Dmf).

f rossii Brignoli, 1977....................Italy [urn:lsid:amnh.org:spidersp:021960]
H. r. Brignoli, 1977f: 79, f. 46 (Df).

mf saccata Zhang, Li & Zheng, 2011....................China [urn:lsid:amnh.org:spidersp:044164]
H. c. Zhang, Li & Zheng, 2011: 16, f. 14A-E, 15A-H, 16A-G (Dmf).

mf sanjuanensis Exline, 1938....................USA, Mexico [urn:lsid:amnh.org:spidersp:021961]
H. s. Exline, 1938: 32, pl. 5, f. 45 (Df).
Hahnistea longipes Chamberlin & Ivie, 1942a: 28, pl. 5, f. 58 (Df).
Hahnistea s. Lehtinen, 1967: 237 (Tf from Hahnia).
H. s. Opell & Beatty, 1976: 426, f. 10, 94-98 (f, Dm, S).

mf schubotzi Strand, 1913....................Central, East Africa [urn:lsid:amnh.org:spidersp:021962]
H. s. Strand, 1913a: 404 (Dmf).
H. macrovulva Strand, 1913a: 405 (Df).
H. rouleti Lessert, 1915b: 493, f. 58-60 (Dmf).
H. macrovulva Roewer, 1955b: 104, f. 39a-b (f).
H. s. Roewer, 1955b: 105, f. 40a-d (mf).
H. kästneri Roewer, 1955b: 107, f. 42 (Df).
H. rouleti Roewer, 1955b: 108, f. 41a-b (mf).
H. kaestneri Roewer, 1955c: 103.
Muizenbergia s. Lehtinen, 1967: 250, f. 360 (Tmf from Hahnia, rejected; S).
Muizenbergia rouleti Lehtinen, 1967: 250 (Tmf from Hahnia, rejected).
Muizenbergia kaestneri Lehtinen, 1967: 250 (Tf from Hahnia, rejected).
H. s. Bosmans, 1986c: 339, f. 1a-c, e, g-h, 2a-d (mf, S).

mf senaria Zhang, Li & Zheng, 2011....................China [urn:lsid:amnh.org:spidersp:044166]
H. s. Zhang, Li & Zheng, 2011: 24, f. 21A-F, 22A-G, 23A-E (Dmf).

mf sexoculata Ponomarev, 2009....................Russia [urn:lsid:amnh.org:spidersp:042942]
H. s. Ponomarev, 2009: 143, f. 4-8 (Dmf).

mf sibirica Marusik, Hippa & Koponen, 1996....................Russia, China [urn:lsid:amnh.org:spidersp:021963]
H. s. Marusik, Hippa & Koponen, 1996: 30, f. 81-84 (Dmf).
H. sibrica Zhang & Zhang, 2003: 52, f. 2A-C (f, lapsus).

f simoni Mello-Leitão, 1919....................Brazil [urn:lsid:amnh.org:spidersp:021964]
H. s. Mello-Leitão, 1919: 478 (Df).

mf sirimoni Benoit, 1978....................Kenya [urn:lsid:amnh.org:spidersp:021965]
H. s. Benoit, 1978e: 617, f. 4A-D (Dmf).

mf spasskyi Denis, 1958....................Afghanistan [urn:lsid:amnh.org:spidersp:021966]
H. s. Denis, 1958b: 118, f. 52-54 (Dmf).
Iberina s. Lehtinen, 1967: 240 (Tmf from Hahnia, rejected).
H. s. Brignoli, 1983c: 505 (rejected transfer).

mf spinata Benoit, 1978....................Kenya [urn:lsid:amnh.org:spidersp:021967]
H. s. Benoit, 1978e: 613, f. 2A-E (Dmf).
H. s. Bosmans & Thijs, 1980: 561, f. 2e (f).

mf submaginii Zhang, Li & Zheng, 2011....................China [urn:lsid:amnh.org:spidersp:044165]
H. s. Zhang, Li & Zheng, 2011: 19, f. 17A-C, 18A-F, 19A-G, 20A-F (Dmf).

mf tabulicola Simon, 1898....................Africa [urn:lsid:amnh.org:spidersp:021968]
H. t. Simon, 1898f: 10 (Dmf).
H. t. Simon, 1898a: 273, f. 282.
H. rouleti annulata Lessert, 1915b: 496 (Df).
H. annulata Roewer, 1955b: 111, f. 43 (f).
H. tubicola Roewer, 1955b: 112, f. 44 (m, lapsus).
Muizenbergia t. Lehtinen, 1967: 250 (Tmf from Hahnia, rejected).
Muizenbergia annulata Lehtinen, 1967: 250 (Tmf from Hahnia, rejected).
H. t. Bosmans, 1981: 204, f. 1-2, 5-6, 9 (mf, S).
H. t. Bosmans, 1982b: 680, f. 8 (f).

mf tatei (Gertsch, 1934)....................Venezuela [urn:lsid:amnh.org:spidersp:021969]
Bigois t. Gertsch, 1934c: 13, f. 5-8, 27-28 (Dmf).
Unzickeria t. Lehtinen, 1967: 273 (Tmf from Bigois=Alistra).

mf thorntoni Brignoli, 1982....................China, Hong Kong, Laos [urn:lsid:amnh.org:spidersp:021970]
H. thortoni Brignoli, 1982b: 346 (Dm; N.B.: f. 13-14 apparently based on misidentified paratype, =H. zhejiangensis).
H. yueluensis Yin & Wang, 1983: 143, f. 8-10 (Df; not m, f. 11-12, =H. zhejiangensis) [urn:lsid:amnh.org:spidersp:021980].
H. maginii Song, 1987: 199, f. 157 (f, misidentified).
H. yueluensis Song, 1987: 202, f. 161A-B (f, not m, f. 161C-D, =H. zhejiangensis).
H. flagellifera Zhu, Chen & Sha, 1989: 148, f. 1-6 (Dmf) [urn:lsid:amnh.org:spidersp:021912].
H. flagellifera Chen & Gao, 1990: 119, f. 149a-f (mf).
H. yueluensis Chen & Gao, 1990: 121, f. 150a (f, not m, f. 150b-c, =H. zhejiangensis).
H. flagellifera Song, Zhu & Chen, 1999: 361, f. 210I-J, 211J-K (mf).
H. maginii Song, Zhu & Chen, 1999: 361, f. 210Q-R (f, misidentified).
H. yueluensis Song, Zhu & Chen, 1999: 362, f. 211B-C (f, not f. 212E-F, =H. zhejiangensis).
H. flagellifera Zhang, Li & Zheng, 2011: 9, f. 6A-C, 7A-D, 8A-I, 9A-G (mf).
H. yueluensis Yin et al., 2012: 960, f. 487b-c (f, misidentified).
H. t. Zhang & Zhang, 2013: 525, f. 3A-E, 4A-I, 5A-C, 6A-G, 7A-D (m, Sf).

f tikaderi Brignoli, 1978....................Bhutan [urn:lsid:amnh.org:spidersp:021971]
H. t. Brignoli, 1978k: 48, f. 26 (Df).

f tortuosa Song & Kim, 1991....................China [urn:lsid:amnh.org:spidersp:021972]
H. t. Song & Kim, 1991: 22, f. 14-15 (Df).
H. t. Song, Zhu & Chen, 1999: 361, f. 210W-X (f).

f tuybaana Barrion & Litsinger, 1995....................Philippines [urn:lsid:amnh.org:spidersp:021973]
H. t. Barrion & Litsinger, 1995: 339, f. 201a-i (Df).

f ulyxis Brignoli, 1974....................Greece [urn:lsid:amnh.org:spidersp:021974]
H. u. Brignoli, 1974b: 170, f. 21 (Df).

f upembaensis Bosmans, 1986....................Congo [urn:lsid:amnh.org:spidersp:021975]
H. u. Bosmans, 1986c: 348, f. 6c-d (Df).

mf vangoethemi Benoit, 1978....................Kenya [urn:lsid:amnh.org:spidersp:021976]
H. v. Benoit, 1978e: 621, f. 5F (Df; not f. 5E).
H. v. Bosmans & Thijs, 1980: 562, f. 1c-d, 2c (f, Dm).

mf vanwaerebeki Bosmans, 1987....................Cameroon [urn:lsid:amnh.org:spidersp:021977]
H. v. Bosmans, 1987: 43, f. 1C, 2A-B (Dmf).

m veracruzana Gertsch & Davis, 1940....................Mexico [urn:lsid:amnh.org:spidersp:021978]
H. v. Gertsch & Davis, 1940a: 14, f. 20 (Dm).
Neohahnia v. Lehtinen, 1967: 251 (Tm from Hahnia, rejected).
H. v. Opell & Beatty, 1976: 432, f. 114-115 (m).

mf xinjiangensis Wang & Liang, 1989....................China [urn:lsid:amnh.org:spidersp:021979]
H. x. Wang & Liang, 1989: 52, f. 1-6 (Dmf).
H. x. Hu & Wu, 1989: 185, f. 154.1-6 (mf).
H. x. Song, Zhu & Chen, 1999: 361, f. 210Y, 211A, 212C-D (mf).

mf yakouensis Chen, Yan & Yin, 2009....................China [urn:lsid:amnh.org:spidersp:042844]
H. y. Chen, Yan & Yin, 2009: 66, f. 1-6 (Dmf).

mf zhejiangensis Song & Zheng, 1982....................China, Taiwan, Vietnam [urn:lsid:amnh.org:spidersp:021981]
H. z. Song & Zheng, 1982b: 81, f. 1-4 (Df).
H. thortoni Brignoli, 1982b: 346, f. 13-14 (m, misidentified).
H. z. Yin & Wang, 1983: 141, f. 6-7 (Dm).
H. yueluensis Yin & Wang, 1983: 143, f. 11-12 (m, misidentified).
H. z. Hu, 1984: 210, f. 222.1-3 (f).
H. thortoni Song, 1987: 201, f. 160 (m, misidentified).
H. yueluensis Song, 1987: 202, f. 161C-D (m, misidentified).
H. z. Song, 1987: 203, f. 162 (mf).
H. z. Feng, 1990: 144, f. 119.1-5 (mf).
H. yueluensis Chen & Gao, 1990: 121, f. 150b-c (m, misidentified).
H. z. Chen & Zhang, 1991: 196, f. 196.1-5 (mf).
H. thorntoni Song, Zhu & Chen, 1999: 361, f. 212A-B (m, misidentified).
H. yueluensis Song, Zhu & Chen, 1999: 362, f. 212E-F (m, misidentified).
H. z. Song, Zhu & Chen, 1999: 362, f. 211D-E, 212G-H (mf).
H. z. Chen, Wang & Chen, 2003: 26, f. 1A-F, 2A-B (mf).
H. z. Zhang, Li & Pham, 2013: 350, f. 7A-E (mf).
H. yueluensis Yin et al., 2012: 960, f. 487d-e (m, misidentified).
H. z. Yin et al., 2012: 961, f. 488a-g (mf).
H. z. Zhang & Zhang, 2013: 529, f. 8A-C, 9A-I, 10A-G, 11A-E (mf).

f zodarioides (Simon, 1898)....................South Africa [urn:lsid:amnh.org:spidersp:021982]
Scotussa z. Simon, 1898a: 276, f. 278 (Df).
H. z. Bosmans, 1992: 90, f. 10-11 (f).