The World Spider Catalog, V15 by N. I. Platnick (c) 2000 - 2014 AMNH

Family: GNAPHOSIDAE

Gen. Haplodrassus Chamberlin, 1922 [urn:lsid:amnh.org:spidergen:02429]

N.B.: considered a senior synonym of Tuvadrassus Marusik & Logunov, 1995b: 194 [urn:lsid:amnh.org:spidergen:02493], type Drassodes tegulatus Schenkel, 1963 by Murphy, 2007: x, 9; see note under Aracus Thorell, 1887.

Transferred to other genera:
H. magister Chamberlin, 1933 -- see Urozelotes.
H. viveki Gajbe, 1992 -- see Drassodes.

In synonymy:
H. admes Chamberlin, 1922 = H. bicornis (Emerton, 1909) (Platnick & Shadab, 1975b: 17).
H. altanus Chamberlin, 1933 = H. hiemalis (Emerton, 1909) (Ubick & Roth, 1973a: 4).
H. capnodes (Thorell, 1875) = H. cognatus (Westring, 1861) (Grimm, 1985: 134).
H. carinatus (Strand, 1906, T from Drassodes) = H. dalmatensis (L. Koch, 1866) (Chatzopoulou & Chatzaki, 2009: 358).
H. denotatus (O. P.-Cambridge, 1874, T from Drassodes) = H. dalmatensis (L. Koch, 1866) (Levy, 2004: 24).
H. desertorum (Caporiacco, 1928, T from Pseudodrassus) = H. dalmatensis (L. Koch, 1866) (Levy, 2004: 24).
H. dystactus Chamberlin & Gertsch, 1940 = H. signifer (C. L. Koch, 1839) (Platnick & Shadab, 1975b: 11).
H. ferrumequinum (F. O. P.-Cambridge, 1899, T from Drassodes) = H. signifer (C. L. Koch, 1839) (Platnick & Shadab, 1975b: 11).
H. gridellii Caporiacco, 1949 = H. pugnans (Simon, 1880) (Levy, 2004: 20).
H. insularis Denis, 1962 = H. dalmatensis (L. Koch, 1866) (Schmidt, 1977: 58).
H. lithobius (Roewer, 1928, T from Drassodes) = H. dalmatensis (L. Koch, 1866) (Chatzaki, Thaler & Mylonas, 2002a: 587).
H. magnipalpus Paik & Sohn, 1984 = H. kulczynskii Lohmander, 1942 (Paik & Kang, 1988: 63).
H. maroccanus Denis, 1956 = H. parvicorpus (Roewer, 1951) (Bosmans & Van Keer, 2012a: 11, contra Levy, 2004: 24, who considered it a S of H. dalmatensis).
H. omissus (O. P.-Cambridge, 1872) = H. morosus (O. P.-Cambridge, 1872) (Levy, 2004: 26).
H. palaestinensis (Strand, 1915, T from Drassodes) = H. dalmatensis (L. Koch, 1866) (Levy, 2004: 24).
H. pseudopugnans (Schenkel, 1963, T from Drassodes by Platnick, 1989b: 475) = H. pugnans (Simon, 1880) (Song & Hubert, 1983: 17, sub Drassodes).
H. uncifer Chamberlin, 1936 = H. bicornis (Emerton, 1909) (Platnick & Shadab, 1975b: 17).
H. utus Chamberlin & Ivie, 1946 = H. eunis Chamberlin, 1922 (Platnick & Shadab, 1975b: 20).
H. vignai Di Franco, 1996 = H. invalidus (O. P.-Cambridge, 1872) (Levy, 2004: 31).
H. walteri (Roewer, 1928, T from Scotophaeus) = H. creticus (Roewer, 1928) (Levy, 2004: 28).

Nomina dubia:
H. barberi (Banks, 1902a: 212, pl. 7, f. 7, 11, f, USA, originally in Prosthesima, T to Zelotes by Petrunkevitch, 1911: 148, here by Chamberlin, 1922: 161) -- Platnick & Shadab, 1975b: 5.
H. pacificus (Banks, 1904b: 336, pl. 39, f. 15, m, USA, originally in Zelotes) -- Platnick & Shadab, 1975b: 5.

f acrotirius (Roewer, 1928)....................Crete [urn:lsid:amnh.org:spidersp:026843]
Drassodes a. Roewer, 1928b: 102, pl. 1, f. 8 (Df).
Drassodes a. Chatzaki, Thaler & Mylonas, 2002a: 587, f. 62A-B (f).
H. a. Bosmans et al., 2013: 10 (Tf from Drassodes).

mf aenus Thaler, 1984....................Switzerland, Austria [urn:lsid:amnh.org:spidersp:027318]
H. a. Thaler, 1984a: 189, f. 1-8, 9a-c, g-h (Dmf).
H. aeneus Grimm, 1985: 132, f. 149, 172-173 (mf).
H. aeneus Heimer & Nentwig, 1991: 426, f. 1126 (mf).

f ambalaensis Gajbe, 1992....................India [urn:lsid:amnh.org:spidersp:027319]
H. a. Gajbe, 1992a: 305, f. 6-10 (Df).

mf atarot Levy, 2004....................Israel [urn:lsid:amnh.org:spidersp:040608]
H. a. Levy, 2004: 33, f. 74-75 (Dm).
H. a. Levy, 2009: 5, f. 6-7 (Df).

mf belgeri Ovtsharenko & Marusik, 1988....................Russia [urn:lsid:amnh.org:spidersp:027320]
H. b. Ovtsharenko & Marusik, 1988: 213, f. 45, 47, 50, 53 (Dmf).
H. b. Ovtsharenko & Marusik, 1996: 121, pl. 2, f. 11-12 (mf).

f bengalensis Gajbe, 1992....................India [urn:lsid:amnh.org:spidersp:037337]
H. b. Gajbe, 1992a: 306, f. 11-15 (Df).

mf bicornis (Emerton, 1909)....................USA, Canada [urn:lsid:amnh.org:spidersp:027321]
Drassus b. Emerton, 1909: 218, pl. 9, f. 2 (Dmf).
H. b. Chamberlin, 1922: 161.
H. admes Chamberlin, 1922: 162 (Df).
H. uncifer Chamberlin, 1936a: 18, f. 17-19 (Dmf).
H. b. Levi & Field, 1954: 458, f. 67, 75-76 (mf).
H. b. Platnick & Shadab, 1975b: 16, f. 23-30 (mf, S).
H. b. Platnick & Dondale, 1992: 220, f. 330-333 (mf).
H. b. Paquin & Dupérré, 2003: 78, f. 673-675 (mf).

mf bohemicus Miller & Buchar, 1977....................Czech Rep., Macedonia, Greece, Ukraine, Russia [urn:lsid:amnh.org:spidersp:027322]
H. b. Miller & Buchar, 1977: 163, pl. II, f. 1-6 (Dmf).
H. b. Stefanovska et al., 2008: 37, f. 10-15 (mf).
H. b. Kovblyuk, Kastrygina & Omelko, 2012: 62, f. 1-6, 10-12 (mf).

mf canariensis Schmidt, 1977....................Canary Is. [urn:lsid:amnh.org:spidersp:027323]
H. c. Schmidt, 1977: 58, f. 5-6 (Dmf).

mf caspius Ponomarev & Belosludtsev, 2008....................Russia, Kazakhstan, Azerbaijan [urn:lsid:amnh.org:spidersp:042161]
H. c. Ponomarev & Belosludtsev, in Ponomarev, Belosludtsev & Dvadnenko, 2008: 164, f. 7-8 (Df).
H. c. Kovblyuk et al., 2013b: 67, f. 1-11 (f, Dm).

mf caucasius Ponomarev & Dvadnenko, 2013....................Russia [urn:lsid:amnh.org:spidersp:046078]
H. c. Ponomarev & Dvadnenko, 2013: 51, f. 1a-b, 2a-b, 3a-c, 4a-b (Dmf).

mf chamberlini Platnick & Shadab, 1975....................North America [urn:lsid:amnh.org:spidersp:027324]
H. c. Platnick & Shadab, 1975b: 27, f. 63-70 (Dmf).
H. c. Platnick & Dondale, 1992: 213, f. 317-321 (mf).

f chotanagpurensis Gajbe, 1987....................India [urn:lsid:amnh.org:spidersp:027325]
H. c. Gajbe, 1987a: 277, f. 1-5 (Df).

mf cognatus (Westring, 1861)....................Palearctic [urn:lsid:amnh.org:spidersp:027326]
Drassus fuscus C. L. Koch, 1843: 123, f. 837 (f, misidentified).
Drassus c. Westring, 1861: 343 (Df).
Drassus fuscus L. Koch, 1866: 86, pl. 4, f. 58 (f).
Drassus capnodes Thorell, 1875a: 96 (Dm).
Drassus capnodes Thorell, 1875c: 92 (Dm).
Drassus c. Dahl, 1883: 56 (Dm).
Drassus buddebergi Simon, 1887c: 142 (Df).
Drassus c. Chyzer & Kulczyn'ski, 1897: 215, pl. 8, f. 53 (mf).
Drassus capnodes Bösenberg, 1902: 296, pl. 28, f. 435 (f).
Drassus c. Bösenberg, 1902: 297, pl. 28, f. 436 (mf).
Drassodes c. Simon, 1914a: 136, 208, f. 231-233 (mf).
H. c. Reimoser, 1937a: 16, f. 34-35 (mf).
H. capnodes Reimoser, 1937a: 17, f. 38 (m).
H. c. Palmgren, 1943: 95, f.
H. c. Tullgren, 1946: 106, f. 31C, pl. 17, f. 221-224 (mf).
H. c. Braendegaard, 1966: 98, f. 85 (f).
H. c. Miller, 1971: 82, pl. VII, f. 17, pl. XXVII, f. 19 (mf).
H. c. Miller & Buchar, 1977: 168, pl. III, f. 8-10 (mf).
H. c. Grimm, 1985: 134, f. 155, 168-169 (mf, S).
H. c. Izmailova, 1989: 105, f. 88 (f).
H. c. Heimer & Nentwig, 1991: 428, f. 1130 (mf).
H. c. Kamura, 1995b: 126, f. 7-8 (f).
H. c. Roberts, 1998: 109, f. (mf).
H. c. Almquist, 2006: 407, f. 351a-f (mf).
H. c. Kamura, 2007: 97, f. 7-8 (m).
H. c. Kamura, 2009: 485, f. 32-34 (mf).
H. c. Kovblyuk, Kastrygina & Omelko, 2012: 66, f. 15-16 (f).

f cognatus ermolajewi Lohmander, 1942....................Russia [urn:lsid:amnh.org:spidersp:027327]
Drassus c. Ermolajev, 1928b: 98, f. 1 (f).
H. c. e. Lohmander, 1942: 86 (Df).

mf concertor (Simon, 1878)....................France [urn:lsid:amnh.org:spidersp:027328]
Drassus c. Simon, 1878a: 129, pl. 16, f. 24 (Dmf).
Drassodes c. Simon, 1914a: 140, 210, f. 249, 251 (mf).
H. c. Denis, 1957c: 248, f. 1-2 (f).

mf creticus (Roewer, 1928)....................Greece, Crete [urn:lsid:amnh.org:spidersp:026876]
Drassodes c. Roewer, 1928b: 104, pl. 1, f. 9 (Dm).
Scotophaeus walteri Roewer, 1928b: 107, pl. 1, f. 18 (Df).
H. c. Chatzaki, Thaler & Mylonas, 2002a: 585, f. 55-60 (Tm from Drassodes, Df).
H. c. Levy, 2004: 28 (S).

mf dalmatensis (L. Koch, 1866)....................Palearctic, Ethiopia [urn:lsid:amnh.org:spidersp:027329]
Drassus d. L. Koch, 1866: 89, pl. 4, f. 59 (Df).
Drassus minusculus L. Koch, 1866: 110, pl. 5, f. 70 (Df).
Drassus denotatus O. P.-Cambridge, 1874a: 398, pl. 52, f. 24 (Df).
Drassus delinquens O. P.-Cambridge, 1875e: 245, pl. 8, f. 4 (Df).
Drassus minusculus L. Koch, 1876b: 302 (Dm).
Drassus minusculus Chyzer & Kulczyn'ski, 1897: 217, pl. 8, f. 58, 63 (mf).
Drassodes carinatus Strand, 1906b: 609 (Dmf) [urn:lsid:amnh.org:spidersp:026869].
Drassodes d. Simon, 1914a: 137, 209, f. 237-239 (mf).
Drassodes palaestinensis Strand, 1915c: 137 (Dm).
Drassodes lithobius Roewer, 1928b: 101, pl. 1, f. 7 (Df).
Scotophaeus desertorum Caporiacco, 1928b: 86, f. 3 (Df).
Pseudodrassus desertorum Caporiacco, 1935a: 286.
Drassodes d. Miller, 1936b: 262, f. 1-2 (mf).
H. d. Tullgren, 1946: 100, pl. 16, f. 201-203 (mf).
H. d. Miller, 1947: 55, pl. II, f. 3 (f).
Drassodes d. Locket & Millidge, 1951: 103, f. 53C, F, H, 54B (mf).
H. insularis Denis, 1962a: 35, f. 22 (Df).
H. d. Braendegaard, 1966: 99, f. 86 (f).
Pseudodrassus desertorum Denis, 1966e: 106, f. 5 (f).
H. d. Tyschchenko, 1971: 97, f. 176 (m).
H. d. Miller, 1971: 83, pl. VII, f. 5-7 (mf).
H. d. Merrett, 1972: 180, f. 1d (m).
H. d. Locket, Millidge & Merrett, 1974: 6, f. 3D (m).
H. d. Schmidt, 1977: 58 (S).
H. d. Miller & Buchar, 1977: 170, pl. IV, f. 1-3 (mf).
H. d. Grimm, 1985: 138, f. 156, 164-165 (mf).
H. d. Roberts, 1985: 66, f. 24a (mf).
H. d. Heimer & Nentwig, 1991: 428, f. 1129 (mf).
H. d. Noordam, 1992: 6 (mf).
H. d. Roberts, 1995: 106, f. (mf).
H. d. Roberts, 1998: 110, f. (mf).
H. d. Chatzaki, Thaler & Mylonas, 2002a: 587, f. 62 (m, S).
H. d. Levy, 2004: 23, f. 57-61 (mf).
H. d. Almquist, 2006: 408, f. 352a-e (mf).
H. d. Chatzopoulou & Chatzaki, 2009: 358 (S).
H. d. Kovblyuk, Kastrygina & Omelko, 2012: 66, f. 22-24, 28-29 (mf).

mf dalmatensis pictus (Thorell, 1875)....................Spain, Madeira [urn:lsid:amnh.org:spidersp:027330]
Drassus p. Thorell, 1875a: 97 (Dmf).
Drassus p. Thorell, 1875c: 90 (Dmf).
Drassodes minusculus p. Kulczyn'ski, 1899: 349, pl. 6, f. 29 (mf).

mf dentatus Xu & Song, 1987....................China [urn:lsid:amnh.org:spidersp:027331]
H. d. Xu & Song, 1987: 83, f. 1-4 (Dmf).
H. d. Xu, 1991b: 37, f. 15 (f).
H. d. Song, Zhu & Chen, 1999: 451, f. 263B, J (mf).
H. d. Song, Zhu & Zhang, 2004: 132, f. 77A-D (mf).
H. d. Omelko & Marusik, 2012: 344, f. 13-15, 19, 25 (mf).

mf deserticola Schmidt & Krause, 1996....................Canary Is. [urn:lsid:amnh.org:spidersp:027332]
H. d. Schmidt & Krause, 1996: 266, f. 4-5 (Df).
H. d. Wunderlich, 2011: 44, f. 84-88 (f, Dm).
H. d. Kovblyuk, Kastrygina & Omelko, 2012: 74, f. 40-42, 50-51, 66-67 (mf).

mf dixiensis Chamberlin & Woodbury, 1929....................USA [urn:lsid:amnh.org:spidersp:027333]
H. d. Chamberlin & Woodbury, 1929: 134, pl. 1, f. 6 (Df).
H. d. Platnick & Shadab, 1975b: 23, f. 39-46 (f, Dm).

f dumdumensis Tikader, 1982....................India [urn:lsid:amnh.org:spidersp:027334]
H. d. Tikader, 1982b: 468, f. 391-393 (Df; probably misplaced).

mf eunis Chamberlin, 1922....................USA, Canada, Alaska [urn:lsid:amnh.org:spidersp:027335]
H. e. Chamberlin, 1922: 162 (Df).
H. utus Chamberlin & Ivie, 1946: 8, f. 9-11 (Dmf).
H. e. Platnick & Shadab, 1975b: 20, f. 31-38 (f, Sm).
H. e. Platnick & Dondale, 1992: 222, f. 334-337 (mf).
H. e. Paquin & Dupérré, 2003: 78, f. 679-681 (mf).

f grazianoi Caporiacco, 1948....................Rhodes [urn:lsid:amnh.org:spidersp:027336]
H. g. Caporiacco, 1948c: 59, f. 10 (Df).

mf hatsushibai Kamura, 2007....................Japan [urn:lsid:amnh.org:spidersp:040609]
H. montanus Chikuni, 1989b: 121, f. 17 (mf, misidentified).
H. h. Kamura, 2007: 100, f. 23-31 (Dmf).
H. h. Kamura, 2009: 487, f. 43-45 (mf).
H. h. Omelko & Marusik, 2012: 344, f. 5-6, 10-12, 18, 23-24 (mf).

mf hiemalis (Emerton, 1909) *....................Holarctic [urn:lsid:amnh.org:spidersp:027338]
Drassus h. Emerton, 1909: 218, pl. 9, f. 1 (Dmf).
H. h. Chamberlin, 1922: 161.
H. altanus Chamberlin, 1933a: 5, f. 8-10 (Dmf).
H. h. Kaston, 1948: 350, f. 1173-1175, 1187 (mf).
H. h. Ubick & Roth, 1973a: 4 (S).
H. h. Platnick & Shadab, 1975b: 7, f. 1-8 (mf).
H. h. Ovtsharenko & Marusik, 1988: 211, f. 54-55, 61 (mf).
H. h. Platnick & Dondale, 1992: 217, f. 326-329 (mf).
H. h. Marusik & Logunov, 1995b: 192, f. 71-75 (mf).
H. h. Ovtsharenko & Marusik, 1996: 120, pl. 2, f. 3-4 (mf).
H. h. Paquin & Dupérré, 2003: 78, f. 676-678 (mf).
H. h. Kamura, 2007: 98, f. 13-16, 21-22 (mf).
H. h. Murphy, 2007: 58, f. 480-481 (mf).
H. h. Omelko & Marusik, 2012: 340, f. 25 (m).

mf huarong Yin & Bao, 2012....................China [urn:lsid:amnh.org:spidersp:046214]
H. h. Yin & Bao, in Yin et al., 2012: 1174, f. 625a-h (Dmf).

f hunanensis Yin & Bao, 2012....................China [urn:lsid:amnh.org:spidersp:046215]
H. h. Yin & Bao, in Yin et al., 2012: 1175, f. 626a-e (Df).

mf invalidus (O. P.-Cambridge, 1872)....................Spain, Corsica, Sicily, Italy, Turkey, Israel, Azerbaijan [urn:lsid:amnh.org:spidersp:027339]
Drassus i. O. P.-Cambridge, 1872a: 237, pl. 15, f. 14 (Dm).
Drassus i. Simon, 1878a: 135 (Df).
Drassodes i. Simon, 1914a: 138, 210, f. 243 (m, not f, =H. mediterraneus).
H. i. Roewer, 1955c: 399.
H. vignai Di Franco, 1996: 173, f. 1-4 (Dmf).
H. i. Levy, 2004: 31, f. 70-73 (mf, S).
H. i. Kovblyuk, Kastrygina & Omelko, 2012: 68, f. 32-34 (m).

mf isaevi Ponomarev & Tsvetkov, 2006....................Greece, Ukraine, Russia, Kazakhstan [urn:lsid:amnh.org:spidersp:041362]
H. i. Ponomarev & Tsvetkov, 2006: 9, f. 12-14 (Dmf).
H. i. Piterkina & Ovtsharenko, 2007: 1426, f. 1.1-6 (mf).
H. i. Kovblyuk, Kastrygina & Omelko, 2012: 70, f. 25-27, 30-31 (mf).

f jacobi Gajbe, 1992....................India [urn:lsid:amnh.org:spidersp:027340]
H. j. Gajbe, 1992b: 315, f. 9-13 (Df).

mf kanenoi Kamura, 1995....................Japan [urn:lsid:amnh.org:spidersp:027341]
H. k. Kamura, 1995b: 123, f. 1-6 (Dmf).
H. k. Kamura, 2007: 97, f. 19-20 (m).
H. k. Kamura, 2009: 485, f. 39-42 (mf).

mf kulczynskii Lohmander, 1942....................Palearctic [urn:lsid:amnh.org:spidersp:027342]
Drassus microps Chyzer & Kulczyn'ski, 1897: 217, pl. 8, f. 57 (mf, misidentified).
Drassus microps Bösenberg, 1902: 300, pl. 28, f. 64, 442 (mf, misidentified).
Drassodes microps Lessert, 1910b: 54 (misidentified).
Drassodes microps Simon, 1914a: 138, 209, f. 240-241 (mf, misidentified).
Drassodes microps Reimoser, 1937a: 18, f. 41-42 (mf, misidentified).
H. k. Lohmander, 1942: 101 (Dmf).
H. microps Denis, 1951a: 104, f. 1-2 (m, misidentified).
H. microps Buchar & Zdárek, 1960: 89, f. 1A-B (m, misidentified).
H. microps Rosca, 1968: 86, f. 19a-b (mf, misidentified).
H. microps Tyschchenko, 1971: 97, f. 177 (m, misidentified).
H. k. Miller, 1971: 82, pl. VII, f. 14-16 (mf).
H. k. Miller & Buchar, 1977: 170, pl. IV, f. 7-10 (mf).
H. magnipalpus Paik & Sohn, 1984: 106, f. 1-13 (Dmf).
H. k. Grimm, 1985: 141, f. 152, 162-163 (mf).
H. k. Paik & Kang, 1988: 63 (S).
H. k. Heimer & Nentwig, 1991: 426, f. 1124 (mf).
H. k. Esyunin & Efimik, 1995: 83, f. 22 (f).
H. k. Roberts, 1998: 111, f. (mf).
H. k. Namkung, 2002: 471, f. 37.12a-c (mf).
H. k. Namkung, 2003: 474, f. 37.12a-c (mf).
H. k. Song, Zhu & Zhang, 2004: 133, f. 78A-F (f).
H. k. Jung et al., 2005: 181, f. 64, 89, 127 (mf).
H. k. Marusik et al., 2007: 43, f. 5-10 (mf).
H. k. Kovblyuk, Kastrygina & Omelko, 2012: 73, f. 35-39 (mf).
H. k. Omelko & Marusik, 2012: 340, f. 28 (m).

m lilliputanus Levy, 2004....................Israel [urn:lsid:amnh.org:spidersp:040610]
H. l. Levy, 2004: 34, f. 76-77 (Dm).

mf macellinus (Thorell, 1871)....................Western Mediterranean [urn:lsid:amnh.org:spidersp:027343]
Drassus m. Thorell, 1871a: 185 (Dmf).
Drassus nyctelinus Thorell, 1871a: 185 (Df).
Drassus occidentalis Thorell, 1871a: 184 (Df).
Drassus typhon Simon, 1878a: 128, pl. 16, f. 20 (Dmf).
Drassodes m. Simon, 1914a: 139, 209, f. 248 (mf).

mf macellinus hebes (O. P.-Cambridge, 1874)....................France, Corsica, Italy, Sicily [urn:lsid:amnh.org:spidersp:027344]
Drassus h. O. P.-Cambridge, 1874a: 371, pl. 51, f. 15 (Dm).
Drassus vicarius Simon, 1878a: 132 (Df).
Drassodes m. h. Simon, 1914a: 140, 209, f. 248 (mf).

mf maculatus (Banks, 1904)....................USA, Mexico [urn:lsid:amnh.org:spidersp:027345]
Zelotes maculata Banks, 1904b: 336, pl. 40, f. 38 (Dm).
H. m. Chamberlin, 1922: 161.
H. m. Platnick & Shadab, 1975b: 26, f. 55-62 (m, Df).

mf mayumiae Kamura, 2007....................Japan [urn:lsid:amnh.org:spidersp:040611]
H. m. Kamura, 2007: 98, f. 9-12, 17-18 (Dmf).
H. m. Kamura, 2009: 485, f. 35-38 (mf).

mf mediterraneus Levy, 2004....................Turkey, Israel [urn:lsid:amnh.org:spidersp:040612]
Drassodes invalidus Simon, 1914a: 138, 210, f. 244 (f, misidentified).
H. m. Levy, 2004: 28, f. 66-69 (Dmf).
H. m. Seyyar, 2011: 1246, f. 1-6 (mf).

mf mimus Chamberlin, 1922....................USA [urn:lsid:amnh.org:spidersp:027347]
H. m. Chamberlin, 1922: 161 (Df).
H. m. Platnick & Shadab, 1975b: 24, f. 47-54 (f, Dm).

mf minor (O. P.-Cambridge, 1879)....................Europe, Ukraine, Russia [urn:lsid:amnh.org:spidersp:027348]
Drassus m. O. P.-Cambridge, 1879e: 192, pl. 12, f. 1 (Df).
Drassus m. Chyzer & Kulczyn'ski, 1897: 219, pl. 8, f. 55, 65 (f, Dm).
Drassus m. Bösenberg, 1902: 300, pl. 28, f. 443 (mf).
Drassodes m. Simon, 1914a: 138, f. 242 (f).
H. m. Reimoser, 1937a: 18, f. 39-40 (mf).
Drassodes m. Locket & Millidge, 1951: 103, f. 54C (mf).
Drassodes m. Locket & Millidge, 1957: 481, f. 1A-B (m).
Drassodes m. Cooke, 1963: 156, f. 4-5 (m).
H. m. Miller, 1971: 82, pl. VI, f. 24 (m).
H. m. Locket, Millidge & Merrett, 1974: 7, f. 2C-D (m).
H. m. Miller & Buchar, 1977: 170, pl. IV, f. 4-6 (m).
H. m. Grimm, 1985: 144, f. 176-178 (mf).
H. m. Roberts, 1985: 66, f. 24c (mf).
H. m. Heimer & Nentwig, 1991: 426, f. 1123 (mf).
H. m. Roberts, 1995: 107, f. (mf).
H. m. Roberts, 1998: 111, f. (mf).
H. m. Chatzaki, Thaler & Mylonas, 2002a: 589, f. 63 (f).
H. m. Tuneva & Esyunin, 2003a: 232, f. 27-33 (mf).
H. m. Deltshev, 2004: 72, f. 6-8 (m).
H. m. Kovblyuk, Kastrygina & Omelko, 2012: 73, f. 43-49, 52-65 (mf).
H. m. Omelko & Marusik, 2012: 340, f. 27 (m).

mf moderatus (Kulczyn'ski, 1897)....................Palearctic [urn:lsid:amnh.org:spidersp:027349]
Drassus m. Kulczyn'ski, in Chyzer & Kulczyn'ski, 1897: 216, pl. 8, f. 60 (Df).
H. m. Lohmander, 1942: 82, 85 (Dm).
H. m. Palmgren, 1943: 96, f.
H. m. Tullgren, 1946: 104, f. 31A, pl. 17, f. 214-216 (mf).
H. modestus Roewer, 1955c: 400 (lapsus).
H. m. Miller, 1971: 82, pl. VIII, f. 2 (m).
H. m. Wunderlich, 1975b: 42, f. 1-5 (mf).
H. m. Izmailova, 1977: 71, f. 3 (f).
H. m. Izmailova, 1978a: 10, f. b (f).
H. m. Grimm, 1985: 145, f. 154, 174-175 (mf).
H. m. Izmailova, 1989: 106, f. 89 (f).
H. m. Heimer & Nentwig, 1991: 428, f. 1132 (mf).
H. m. Marusik & Logunov, 1995b: 192, f. 76-80 (mf).
H. m. Roberts, 1998: 109, f. (mf).
H. m. Song, Zhu & Zhang, 2004: 134, f. 79A-C (f).
H. m. Almquist, 2006: 409, f. 353a-g (mf).
H. m. Marusik & Kovblyuk, 2011: 156, f. 19 (f).

mf montanus Paik & Sohn, 1984....................Russia, China, Korea [urn:lsid:amnh.org:spidersp:027350]
H. m. Paik & Sohn, 1984: 107, f. 14-26 (Dmf).
H. m. Namkung, 2002: 470, f. 37.11a-b (mf).
H. m. Namkung, 2003: 473, f. 37.11a-b (mf).
H. m. Jung et al., 2005: 182, f. 65, 128 (mf).
H. m. Omelko & Marusik, 2012: 345, f. 1-4, 7-9, 16-17, 20-22 (mf).
H. m. Yin et al., 2012: 1177, f. 627a-f (mf).

mf morosus (O. P.-Cambridge, 1872)....................Greece, Turkey, Israel, Karakorum [urn:lsid:amnh.org:spidersp:026954]
Drassus m. O. P.-Cambridge, 1872a: 232, pl. 15, f. 9 (Dm).
Drassus omissus O. P.-Cambridge, 1872a: 239, pl. 15 (Df).
H. omissus Caporiacco, 1934a: 131.
H. m. Levy, 2004: 26, f. 62-65 (Tm from Drassodes, Sf).

f nojimai Kamura, 2007....................Japan [urn:lsid:amnh.org:spidersp:040613]
H. n. Kamura, 2007: 102, f. 32-33 (Df).
H. n. Kamura, 2009: 487, f. 46 (f).

mf ovtchinnikovi Ponomarev, 2008....................Turkey, Kazakhstan [urn:lsid:amnh.org:spidersp:041586]
H. o. Ponomarev, 2008a: 57, f. 26-28 (Dmf).
H. o. Kovblyuk et al., 2009: 177, f. 42-50 (mf).

f paramecus Zhang, Song & Zhu, 2001....................China [urn:lsid:amnh.org:spidersp:038063]
H. p. Zhang, Song & Zhu, 2001: 53, f. 2A-C (Df).
H. p. Song, Zhu & Zhang, 2004: 136, f. 80A-C (f).

f pargongsanensis Paik, 1992....................Korea [urn:lsid:amnh.org:spidersp:027352]
H. p. Paik, 1992j: 88, f. 9-14 (Df; N.B.: erroneously spelled H. pargongensis in legend).
H. p. Jung et al., 2005: 183, f. 23, 66, 90 (f).

mf parvicorpus (Roewer, 1951)....................Mallorca, Morocco [urn:lsid:amnh.org:spidersp:026966]
Drassus parvulus L. Koch, 1882: 632, pl. 1, f. 10-11 (Dm, preoccupied).
H. maroccanus Denis, 1956a: 196, f. 4-6 (Dmf).
Drassodes p. Roewer, 1951: 443 (replacement name for Drassus parvulus)
H. p. Bosmans & Van Keer, 2012a: 11, f. 27-28 (Tm from Drassodes, Sf).

mf ponomarevi Kovblyuk & Seyyar, 2009....................Turkey [urn:lsid:amnh.org:spidersp:042878]
H. p. Kovblyuk & Seyyar, in Kovblyuk et al., 2009: 177, f. 51-62 (Dmf).

mf pseudosignifer Marusik, Hippa & Koponen, 1996....................Ukraine, Russia [urn:lsid:amnh.org:spidersp:027353]
H. p. Marusik, Hippa & Koponen, 1996: 26, f. 63-65, 69 (Dmf).
H. p. Kovblyuk, Kastrygina & Omelko, 2012: 78, f. 68-70, 75-76 (mf).

mf pugnans (Simon, 1880)....................Palearctic [urn:lsid:amnh.org:spidersp:027354]
Drassus p. Simon, 1880b: 118, pl. 3, f. 20-21 (Dmf).
H. gridellii Caporiacco, 1949b: 117, f. 3 (Df).
Drassodes p. Schenkel, 1963: 43, f. 20a-c (mf).
Drassodes pseudopugnans Schenkel, 1963: 45, f. 21a-c (Dm).
Drassodes p. Song & Hubert, 1983: 17, f. 50-54 (mf, S).
H. signifer Hayashi, 1984: 13, f. 12-26 (mf, misidentified).
H. p. Yaginuma, 1986a: 193, f. 108.1 (Tmf from Drassodes).
H. p. Song, 1987: 340, f. 295 (mf).
Drassodes p. Zhang, 1987: 175, f. 147.1-5 (mf).
H. p. Ovtsharenko & Marusik, 1988: 213, f. 56-58 (mf).
H. p. Chikuni, 1989b: 121, f. 16 (mf).
H. p. Ovtsharenko & Marusik, 1996: 120, pl. 2, f. 5-6 (mf).
H. p. Song, Zhu & Chen, 1999: 451, f. 263C, K (mf).
H. p. Hu, 2001: 255, f. 144.1-5 (mf).
H. p. Song, Zhu & Chen, 2001: 345, f. 223A-D (mf).
H. p. Levy, 2004: 20, f. 53-56 (mf, S).
H. p. Song, Zhu & Zhang, 2004: 137, f. 81A-J (mf).
H. p. Kamura, 2007: 95, f. 1-6 (mf).
H. p. Kamura, 2009: 485, f. 29-31 (mf).
H. p. Kovblyuk, Kastrygina & Omelko, 2012: 64, f. 7-9, 13-14 (mf).

f reginae Schmidt & Krause, 1998....................Cape Verde Is. [urn:lsid:amnh.org:spidersp:027355]
H. r. Schmidt & Krause, 1998: 421, f. 4 (Df).

mf rufus (Savelyeva, 1972)....................Kazakhstan [urn:lsid:amnh.org:spidersp:027356]
Aphantaulax rufa Savelyeva, 1972a: 1238, f. 5-8 (Dmf).
Aphantaulax r. Brignoli, 1983c: 565.
H. r. Ovtsharenko & Savelyeva, 1992: 132 (Tmf from Aphantaulax).

m rugosus Tuneva, 2005....................Kazakhstan [urn:lsid:amnh.org:spidersp:040614]
H. r. Tuneva, 2005: 322, f. 6-9 (Dm).

mf sataraensis Tikader & Gajbe, 1977....................India [urn:lsid:amnh.org:spidersp:027357]
H. s. Tikader & Gajbe, 1977c: 70, f. 4A-E (Dmf).
H. s. Tikader, 1982b: 466, f. 385-390 (mf).
H. s. Gajbe, 2007: 479, f. 167-172 (mf).

mf seditiosus (Caporiacco, 1928)....................Libya [urn:lsid:amnh.org:spidersp:027358]
Drassodes s. Caporiacco, 1928b: 83, f. 1 (Df).
H. s. Caporiacco, 1934b: 9, f. 3 (Dm).

mf severus (C. L. Koch, 1839)....................Mediterranean [urn:lsid:amnh.org:spidersp:027359]
Drassus s. C. L. Koch, 1839a: 22, f. 446 (Df).
Clubiona s. Walckenaer, 1841: 479.
Drassus s. C. L. Koch, 1843: 126, f. 838 (Dm).
Drassus similis L. Koch, 1866: 103, pl. 4, f. 67 (Dmf).
Drassus s. L. Koch, 1866: 108, pl. 5, f. 69 (f).
Drassodes s. Simon, 1893a: 361, f. 322 (m).
Drassodes s. Simon, 1914a: 136, 210, f. 234-236 (mf).

mf signifer (C. L. Koch, 1839)....................Holarctic [urn:lsid:amnh.org:spidersp:027360]
Drassus s. C. L. Koch, 1839a: 31, f. 452 (Df).
Drassus troglodytes C. L. Koch, 1839a: 35, f. 455-456 (Dmf).
Clubiona troglodytes Walckenaer, 1841: 480.
Drassus clavator O. P.-Cambridge, 1860: 171 (Dmf).
Drassus clavator Blackwall, 1861a: 109, pl. 6, f. 66 (m).
Drassus troglodytes L. Koch, 1866: 116, pl. 5, f. 73-74 (mf).
Drassus troglodytes Menge, 1875: 378, pl. 65, f. 216 (mf).
Drassus robustus Emerton, 1890: 179, pl. 4, f. 8 (Dmf).
Drassus mysticus O. P.-Cambridge, 1894b: 104 (D).
Teminius nigriceps Banks, 1895b: 421 (Df).
Drassus placidus Banks, 1896e: 63 (Df).
Drassus troglodytes Becker, 1896: 257, pl. 16, f. 13 (mf).
Drassus troglodytes Chyzer & Kulczyn'ski, 1897: 217, pl. 8, f. 56, 61 (mf).
Drassodes ferrum-equinum F. O. P.-Cambridge, 1899a: 60, pl. 4, f. 12 (Df).
Prosthesima decepta Banks, 1900c: 531 (Df).
Drassus troglodytes Bösenberg, 1902: 298, pl. 28, f. 439 (mf).
Drassodes robustus Bryant, 1908: 7.
Drassodes troglodytes Lessert, 1910b: 52.
Zelotes decepta Petrunkevitch, 1911: 149.
Syrisca nigriceps Petrunkevitch, 1911: 514.
Drassodes s. Comstock, 1912: 313, f. 303-304.
Drassodes troglodytes Simon, 1914a: 122, 140, 209, f. 249-250 (mf).
Drassodes beaufortensis Strand, 1916b: 95 (provisional name only).
H. s. Chamberlin, 1922: 163.
Drassus troglodytes Spassky, 1925: 35, f. 4 (f).
H. s. Reimoser, 1937a: 17, f. 36-37 (mf).
H. dystactus Chamberlin & Gertsch, 1940: 8, f. 6, 9-10 (Dmf).
H. s. Comstock, 1940: 326, f. 303 (f).
H. s. Palmgren, 1943: 95, f.
H. s. Tullgren, 1946: 98, f. 30A, pl. 16, f. 197-200 (mf).
Drassodes s. Braendegaard, 1946: 55, g. 35-36 (mf).
H. s. Kaston, 1948: 350, f. 1170-1172, 1186 (mf).
Drassodes s. Locket & Millidge, 1951: 101, f. 53A, D, G, 54A (mf).
H. s. Braendegaard, 1966: 94, f. 81-82 (mf).
H. s. Tyschchenko, 1971: 97, f. 175 (m).
H. s. Miller, 1971: 82, pl. VII, f. 18, pl. XXVII, f. 20-21 (mf).
H. s. Platnick & Shadab, 1975b: 11, f. 11-22 (mf, S).
H. s. Miller & Buchar, 1977: 168, pl. II, f. 7-10 (mf).
H. s. Paik, 1978e: 417, f. 188.1-5 (f).
Drassodes s. Jia & Zhu, 1983: 167, f. a-e (mf).
H. s. Thaler, 1984a: 189, f. 9d-f, i (m).
H. s. Grimm, 1985: 146, f. 146-148, 170-171 (mf).
H. s. Roberts, 1985: 66, f. 23g (mf).
H. s. Heiss & Allen, 1986: 57, f. 95-96 (mf).
H. aeneus Hu & Li, 1987b: 301, f. 32.1-4 (mf, misidentified).
H. s. Izmailova, 1989: 107, f. 90 (m).
H. s. Heimer & Nentwig, 1991: 426, f. 1125 (mf).
H. s. Platnick & Dondale, 1992: 215, f. 322-325 (mf).
H. s. Paik, 1992j: 89, text-f. 1A-B (f).
H. s. Noordam, 1992: 6 (mf).
H. s. Roberts, 1995: 106, f. (mf).
H. s. Marusik, Hippa & Koponen, 1996: 26, f. 66-68, 70 (mf).
H. s. Ovtsharenko & Marusik, 1996: 121, pl. 2, f. 7-8 (mf).
H. s. Agnarsson, 1996: 31, f. 11A-C (mf).
H. s. Mcheidze, 1997: 114, f. 157-160 (mf).
H. s. Bellmann, 1997: 170, f. (f).
H. s. Roberts, 1998: 109, f. (mf).
Drassodes s. Hu, 2001: 240, f. 132.1-2 (mf).
H. aeneus Hu, 2001: 255, f. 143.1-4 (mf, misidentified).
H. s. Chatzaki, Thaler & Mylonas, 2002a: 587, f. 61 (m).
H. s. Paquin & Dupérré, 2003: 78, f. 682-684 (mf).
H. s. Levy, 2004: 19, f. 46-52 (mf).
H. s. Song, Zhu & Zhang, 2004: 139, f. 82A-I (mf).
H. s. Almquist, 2006: 411, f. 354a-i (mf).
H. s. Kovblyuk, Kastrygina & Omelko, 2012: 81, f. 71-74, 77-78 (mf).
H. s. Omelko & Marusik, 2012: 340, f. 26 (m).

mf silvestris (Blackwall, 1833)....................Palearctic [urn:lsid:amnh.org:spidersp:027361]
Drassus sylvestris Blackwall, 1833: 440 (Df; emendation now amply protected by usage).
Drassus sylvestris Blackwall, 1861a: 113, pl. 6, f. 68 (f, Dm).
Drassus infuscatus Westring, 1861: 347 (Df).
Drassus infuscatus L. Koch, 1866: 99, pl. 4, f. 64 (f, Dm).
Drassus criminalis O. P.-Cambridge, 1875e: 244, pl. 8, f. 3 (Df).
Drassus infuscatus Menge, 1875: 381, pl. 66, f. 218 (mf).
Drassus infuscatus Becker, 1896: 256, pl. 16, f. 12 (mf).
Drassus s. Chyzer & Kulczyn'ski, 1897: 216, pl. 8, f. 45, 54 (mf).
Drassus infuscatus Bösenberg, 1902: 299, pl. 28, f. 440 (mf).
Drassodes s. Simon, 1914a: 135, 208, f. 228-230 (mf).
H. s. Reimoser, 1937a: 15, f. 32-33 (mf).
H. s. Tullgren, 1946: 105, f. 31B, pl. 17, f. 217-220 (mf).
Drassodes s. Locket & Millidge, 1951: 103, f. 53B, E, 54D (mf).
H. s. Braendegaard, 1966: 96, f. 83-84 (mf).
H. s. Miller, 1971: 82, pl. VII, f. 2-4 (mf).
H. s. Miller & Buchar, 1977: 168, pl. III, f. 5-7 (mf).
H. s. Grimm, 1985: 150, f. 18b, 153, 166-167 (mf).
H. s. Roberts, 1985: 66, f. 24b (mf).
H. s. Izmailova, 1989: 107, f. 91 (f).
H. s. Heimer & Nentwig, 1991: 428, f. 1131 (mf).
H. s. Noordam, 1992: 6 (mf).
H. s. Roberts, 1995: 107, f. (mf).
H. s. Roberts, 1998: 110, f. (mf).
H. s. Almquist, 2006: 411, f. 355a-f (mf).
H. s. Seyyar, 2011: 1248, f. 7-9 (m).
H. s. Kovblyuk, Kastrygina & Omelko, 2012: 66, f. 17-21 (mf).
H. s. Ponomarev & Dvadnenko, 2013: 52, f. 1c-d, 2c-d, 3d (mf).

mf soerenseni (Strand, 1900)....................Palearctic [urn:lsid:amnh.org:spidersp:027362]
Drassus s. Strand, 1900a: 98, f. (Df).
H. lapponicus Holm, 1939b: 4, f. 1a-d, 2 (Dmf).
H. s. Lohmander, 1942: 101.
H. s. Tullgren, 1942: 221, 230, pl. 1, f. 3 (mf).
H. s. Palmgren, 1943: 97, f.
H. s. Tullgren, 1946: 102, f. 30C, pl. 17, f. 209-213 (mf).
H. sörenseni Locket & Millidge, 1953: 406, f. 243A-C (mf).
H. s. Miller, 1971: 84, pl. VII, f. 8-10 (mf).
H. sörenseni Merrett, 1972: 180, f. 1e (m).
H. s. Locket, Millidge & Merrett, 1974: 6, f. 3E (m).
H. s. Miller & Buchar, 1977: 168, pl. II, f. 11-12, pl. III, f. 1 (mf).
H. s. Grimm, 1985: 153, f. 151, 160-161 (mf).
H. s. Roberts, 1985: 66, f. 24d (mf).
H. s. Ovtsharenko & Marusik, 1988: 214, f. 48, 51-52 (mf).
H. s. Heimer & Nentwig, 1991: 428, f. 1127 (mf).
H. s. Roberts, 1995: 107, f. (mf).
H. s. Ovtsharenko & Marusik, 1996: 121, pl. 2, f. 9-10 (mf).
H. s. Roberts, 1998: 111, f. (mf).
H. s. Song, Zhu & Zhang, 2004: 141, f. 83A-C (f).
H. s. Almquist, 2006: 412, f. 356a-e (mf).

f stuxbergi (L. Koch, 1879)....................Russia [urn:lsid:amnh.org:spidersp:027363]
Drassus s. L. Koch, 1879c: 87, pl. 3, f. 6 (Df).
Drassodes s. Kulczyn'ski, 1916: 2, pl. 1, f. 1 (f).
H. s. Holm, 1973: 103 (Tf from Drassodes).

mf taepaikensis Paik, 1992....................Russia, Korea [urn:lsid:amnh.org:spidersp:027364]
H. t. Paik, 1992j: 86, f. 1-8 (Df).
H. t. Jung et al., 2005: 184, f. 25, 67, 91 (f).
H. t. Marusik et al., 2007: 43, f. 1-4, 11-18 (f, Dm).
H. t. Marusik & Kovblyuk, 2011: 156, f. 12, 16 (mf).

mf taibo (Chamberlin, 1919)....................USA [urn:lsid:amnh.org:spidersp:027365]
Zelotes t. Chamberlin, 1919b: 6, pl. 2, f. 5 (Df).
H. t. Chamberlin, 1922: 161.
H. t. Platnick & Shadab, 1975b: 29, f. 71-74 (f, Dm).

mf tegulatus (Schenkel, 1963)....................Russia, China [urn:lsid:amnh.org:spidersp:028043]
Drassodes t. Schenkel, 1963: 40, f. 18 (Df).
Tuvadrassus t. Marusik & Logunov, 1995b: 194, f. 83-87 (Tf from Drassodes, Dm).
Tuvadrassus t. Song, Zhu & Zhang, 2004: 238, f. 139A-D (mf).
H. t. Murphy, 2007: 58, f. 486-487 (f).
H. t. Omelko & Marusik, 2012: 341, f. 29 (m).

mf tehriensis Tikader & Gajbe, 1977....................India [urn:lsid:amnh.org:spidersp:027366]
H. t. Tikader & Gajbe, 1977c: 68, f. 3A-E (Dmf).
H. t. Tikader, 1982b: 464, f. 379-384 (mf).

mf umbratilis (L. Koch, 1866)....................Europe to Kazakhstan [urn:lsid:amnh.org:spidersp:027367]
Drassus u. L. Koch, 1866: 113, pl. 5, f. 71 (Df).
Drassus microps Menge, 1875: 380, pl. 65, f. 217 (Dmf).
Drassus u. Chyzer & Kulczyn'ski, 1897: 217, pl. 8, f. 59, 62 (mf).
Drassus u. Bösenberg, 1902: 299, pl. 28, f. 441 (mf).
Drassodes u. Simon, 1914a: 138, 209, f. 240-247 (mf).
Drassus u. Spassky, 1925: 35, f. 24 (m).
H. u. Reimoser, 1937a: 19, f. 43-44 (mf).
H. u. Holm, 1939b: 5, f. 1e (m).
H. u. Palmgren, 1943: 96, f.
H. u. Tullgren, 1946: 101, f. 30B, pl. 16, f. 204-208 (mf).
H. u. Braendegaard, 1966: 100, f. 87 (f).
H. u. Rosca, 1968: 86, f. 20a-b (mf).
H. u. Tyschchenko, 1971: 97, f. 178 (m).
H. u. Miller, 1971: 84, pl. VII, f. 11-13 (mf).
H. u. Merrett, 1972: 179, f. 1a-c (mf).
H. u. Locket, Millidge & Merrett, 1974: 7, f. 3A-C (mf).
H. u. Miller & Buchar, 1977: 168, pl. III, f. 2-4 (mf).
H. u. Grimm, 1985: 156, f. 150, 158-159 (mf).
H. u. Roberts, 1985: 66, f. 25a (mf).
H. u. Heimer & Nentwig, 1991: 428, f. 1128 (mf).
H. u. Roberts, 1995: 108, f. (mf).
H. u. Roberts, 1998: 112, f. (mf).
H. u. Almquist, 2006: 413, f. 357a-f (mf).
H. u. Kovblyuk, Kastrygina & Omelko, 2012: 84, f. 79-83 (mf).

f umbratilis gothicus Lohmander, 1942....................Sweden [urn:lsid:amnh.org:spidersp:027368]
H. u. g. Lohmander, 1942: 101 (Df).

f vastus (Hu, 1989)....................China [urn:lsid:amnh.org:spidersp:028434]
Zelotes v. Hu, 1989: 99, f. 3-4 (Df).
Zelotes v. Hu & Wu, 1989: 295, f. 238.1-2 (f).
H. v. Song, Zhu & Zhang, 2004: 143, f. 84A-B (Tf from Zelotes).