The World Spider Catalog, V15 by N. I. Platnick (c) 2000 - 2014 AMNH

Family: LIPHISTIIDAE

Gen. Heptathela Kishida, 1923 [urn:lsid:amnh.org:spidergen:00001]

N.B.: not a senior synonym of Ryuthela Haupt, 1983 (Haupt, 2003: 91, contra Raven, 1985a: 16); considered a senior synonym of Abcathela Ono, 2000: 149, type H. abca Ono, 1999 and Vinathela Ono, 2000: 150, type H. cucphuongensis Ono, 1999 by Haupt, 2003: 91, and of Songthela Ono, 2000: 150, type H. hangzhouensis Chen, Zhang & Zhu, 1981 [urn:lsid:amnh.org:spidergen:03548], Nanthela Haupt, 2003: 69, type Heptathela tonkinensis (Bristowe, 1933) [urn:lsid:amnh.org:spidergen:03546], and Sinothela Haupt, 2003: 69, type Heptathela sinensis Bishop & Crosby, 1932 by Schwendinger & Ono, 2011: 601 (the latter genus had been placed as a junior S of Songthela in an earlier version of this catalog).

Transferred to other genera:
H. nishihirai Haupt, 1979 -- see Ryuthela.

In synonymy:
H. suoxiyuensis Yin, Tang & Xu, 2003 = H. ciliensis Yin, Tang & Xu, 2003 (Yin et al., 2012: 112).
H. xianensis Zhu & Wang, 1983 = H. schensiensis (Schenkel, 1953) (Song & Haupt, 1984: 447).
H. yuelushanensis Wang & Ye, 1983 = H. hangzhouensis Chen, Zhang & Zhu, 1981 (Platnick, 1989b: 57, sub Heptathela).

Nomen nudum:
H. beihequan Yin, 2001b: 2, f. 2I (f, China).

f abca Ono, 1999....................Vietnam [urn:lsid:amnh.org:spidersp:000001]
H. a. Ono, 1999: 37, f. 2-6, 12 (Df).
Abcathela a. Ono, 2000: 150.

mf amamiensis Haupt, 1983....................Ryukyu Is. [urn:lsid:amnh.org:spidersp:000002]
H. kimurai a. Haupt, 1983: 283, f. 9b (Df).
H. a. Ono & Nishikawa, 1989: 120, f. 1-7 (f, Dm, elevated to species).
H. kimurai a. Haupt, 2003: 69, f. 43I, K, 49A-B, 52P, 54N-O, 55.1-3, 62G (mf).
H. a. Ono, 2009a: 80, f. 2-3, 41-47 (mf).

mf australis (Ono, 2002)....................Vietnam [urn:lsid:amnh.org:spidersp:037418]
Songthela a. Ono, 2002c: 120, f. 1-8 (Df).
H. a. Schwendinger & Ono, 2011: 602, f. 1-20, 55-58, 67 (f, Dm).

mf bristowei Gertsch, 1967....................China [urn:lsid:amnh.org:spidersp:000003]
H. b. Gertsch, 1967b: 116, f. 1-3 (Df).
H. b. Haupt, 1983: 285, f. 9e (f).
H. b. Yin, Wang & Hu, 1983: 34, f. 6D (Dm).
H. b. Song, Zhu & Chen, 1999: 32, f. 15A (f).
Abcathela b. Ono, 2000: 150.
H. bristowai Yin et al., 2012: 34, f. 3-16a-b (m, lapsus).

mf ciliensis Yin, Tang & Xu, 2003....................China [urn:lsid:amnh.org:spidersp:038355]
H. c. Yin, Tang & Xu, 2003: 1, f. 1.1-13 (Dmf).
H. suoxiyuensis Yin, Tang & Xu, 2003: 3, f. 2.1-11 (Dmf) [urn:lsid:amnh.org:spidersp:038357].
H. c. Yin et al., 2012: 112, f. 1a-m (mf, S).

f cipingensis (Wang, 1989)....................China [urn:lsid:amnh.org:spidersp:000004]
Liphistius c. Wang, 1989a: 30, f. 1-6 (Df)
H. c. Platnick, 1993c: 77 (Tf from Liphistius).
Songthela c. Ono, 2000: 150.

f cucphuongensis Ono, 1999....................Vietnam [urn:lsid:amnh.org:spidersp:000005]
H. c. Ono, 1999: 38, f. 7-9, 17 (Df).
Vinathela c. Ono, 2000: 150.

f goulouensis Yin, 2001....................China [urn:lsid:amnh.org:spidersp:037420]
H. g. Yin, 2001a: 297, f. 1-9 (Df).
H. g. Yin, 2001b: 2, f. 3A-I (f).
H. g. Yin et al., 2012: 114, f. 2a-i (f).

mf hangzhouensis Chen, Zhang & Zhu, 1981....................China [urn:lsid:amnh.org:spidersp:000006]
H. h. Chen, Zhang & Zhu, 1981: 305, f. 1-7 (Dm).
H. h. Haupt, 1983: 285, f. 8c (m).
H. yuelushanensis Wang & Ye, 1983: 146, f. 1-9 (Dmf).
H. bristowei Zhu, 1983: 4 (S, rejected).
H. h. Hu, 1984: 28, f. 27.1-2 (m).
H. h. Song & Haupt, 1984: 448, f. 2c, 2e-f, 3c-d (Df).
H. yuelushanensis Song & Haupt, 1984: 449 (considered a nomen dubium).
H. yuelushanensis Yin et al., 1988: 53, f. IA-H, IIA-G, IIIA-F, IVA-F (mf).
H. h. Platnick, 1989b: 57 (S).
H. h. Feng, 1990: 26, f. 1.1-4 (mf).
H. yuelushanensis Chen & Gao, 1990: 16, f. 19a-d (mf).
H. h. Chen & Zhang, 1991: 30, f. 24.1-8 (mf).
H. h. Song, Zhu & Chen, 1999: 32, f. 15B-C, H-I (mf).
Songthela h. Ono, 2000: 150.
H. Yin, 2001b: 1, f. 1A-F, 2A-H, 4A-E (mf).
H. h. Yoo & Kim, 2002: 27, f. 44-45 (m).
Sinothela h. Haupt, 2003: 71, f. 52L, 62D (mf).
Songthela h. Yin et al., 2012: 119, f. 6a-e (mf).

f heyangensis (Zhu & Wang, 1984)....................China [urn:lsid:amnh.org:spidersp:000007]
Liphistius h. Zhu & Wang, 1984: 251, f. 1-7 (Df).
H. h. Platnick, 1989b: 57 (Tf from Liphistius).
Abcathela h. Ono, 2000: 150.
Sinothela h. Haupt, 2003: 71 (Tf from Heptathela).

mf higoensis Haupt, 1983....................Japan [urn:lsid:amnh.org:spidersp:000008]
H. kimurai h. Haupt, 1983: 283, f. 12a, 13a (Dmf).
H. h. Ono, 1998: 16, f. 5-10 (mf, elevated from subspecies).
H. kimurai h. Haupt, 2003: 69, f. 43G (f).
H. kimurai h. Haupt, 2003: 69, f. 50C-D, 52O, 54I-L, 62K (mf).
H. h. Ono, 2009a: 80, f. 60-65 (mf).

m hongkong Song & Wu, 1997....................Hong Kong [urn:lsid:amnh.org:spidersp:000009]
H. h. Song & Wu, 1997: 1, f. 1A-C (Dm).
H. h. Song, Zhu & Chen, 1999: 33, f. 11A, 15J-K (m).
Vinathela h. Ono, 2000: 150.
Nanthela h. Haupt, 2003: 69, f. 51C-E (Tm from Heptathela).

f hunanensis Song & Haupt, 1984....................China [urn:lsid:amnh.org:spidersp:000010]
H. h. Song & Haupt, 1984: 449, f. 3e (Df).
H. h. Song, Zhu & Chen, 1999: 33, f. 15D (f).
Vinathela h. Ono, 2000: 150.
H. h. Yin et al., 2012: 115, f. 3a-c (f).

mf jianganensis Chen et al., 1988....................China [urn:lsid:amnh.org:spidersp:000012]
H. j. Chen et al., 1988: 78, f. 1-9 (Dmf).
H. j. Chen & Gao, 1990: 14, f. 18a-g (mf).
Abcathela j. Ono, 2000: 150.

mf kanenoi Ono, 1996....................Ryukyu Is. [urn:lsid:amnh.org:spidersp:000013]
H. k. Ono, 1996b: 158, f. 1-5 (Dmf).
H. k. Ono, 2009a: 80, f. 48-52 (mf).

mf kikuyai Ono, 1998....................Japan [urn:lsid:amnh.org:spidersp:000014]
H. k. Ono, 1998: 16, f. 11-16 (Dmf).
H. k. Ono, 2009a: 81, f. 5-6, 75-80 (mf).
H. k. Schwendinger & Ono, 2011: 613, f. 44-45 (m).

mf kimurai (Kishida, 1920) *....................Japan [urn:lsid:amnh.org:spidersp:000015]
Liphistius k. Kishida, 1920b: 362, f. 1 (Dmf).
H. k. Kishida, 1923: 236, f. 1-7 (f).
H. k. Bristowe, 1933a: 1030, f. 8g, 9b.
H. k. Sawaguti & Ozi, 1937: 116, f. 1-5 (m).
H. k. Yaginuma, 1954b: 15, pl. I (m).
H. k. Yaginuma, 1955b: 35, f. 2.1-3 (m).
H. k. Yaginuma, 1960: 19, f. 12 (m).
H. k. Yaginuma, 1971: 19, f. 12 (m).
H. k. Gertsch & Platnick, 1979: 5, f. 13-14.
H. k. Yaginuma, 1979c: 1, f. 1 (m).
H. k. Yaginuma, 1980c: 44, f. 1 (m).
H. k. Haupt, 1983: 283, f. 1b, 7a-b, 9a, 11a, 12c, 13b-c (mf).
H. k. Yoshikura, 1983: 63, f. 1-7 (mf).
H. k. Haupt, 1984: 163, f. 2a (m).
H. k. Yaginuma, 1986a: 1, f. 1.1 (m).
H. k. Yoshikura, 1987: 148, f. 20.3A, 20.14, 35.2A (mf).
H. k. Chikuni, 1989b: 18, f. 1 (m).
H. k. Ono, 1998: 14, f. 1-4 (m).
H. k. Yoo & Kim, 2002: 27, f. 46 (m).
H. k. Haupt, 2003: 69, f. 50A-B, 52M-N, 54A-H, 62L (mf).
H. k. Ono, 2009a: 81, f. 4, 68-74 (mf).
H. k. Schwendinger & Ono, 2011: 614, f. 46-47 (m).

f luotianensis Yin et al., 2002....................China [urn:lsid:amnh.org:spidersp:037422]
H. l. Yin et al., 2002: 18, f. 1-8 (Df).

m mangshan Bao, Yin & Xu, 2003....................China [urn:lsid:amnh.org:spidersp:038356]
H. m. Bao, Yin & Xu, 2003: 459, f. 1-3 (Dm).
H. m. Yin et al., 2012: 116, f. 4a-c (m).

mf nishikawai Ono, 1998....................Japan [urn:lsid:amnh.org:spidersp:000018]
H. n. Ono, 1998: 19, f. 17-18 (Df).
H. n. Ono, 2009a: 83, f. 81-84 (f, Dm).

mf nui Schwendinger & Ono, 2011....................Vietnam [urn:lsid:amnh.org:spidersp:044561]
H. n. Schwendinger & Ono, 2011: 607, f. 21-33, 68 (Dmf).

f sapana (Ono, 2010)....................Vietnam [urn:lsid:amnh.org:spidersp:043283]
Abcathela s. Ono, 2010b: 3, f. 11-16 (Df).

mf schensiensis (Schenkel, 1953)....................China [urn:lsid:amnh.org:spidersp:000021]
Liphistius (Heptathele) sinensis s. Schenkel, 1953b: 1, f. 1a-d (Df).
Liphistius s. Gertsch, 1967b: 115 (elevated to species).
H. sinensis s. Haupt, 1983: 285, f. 9d (Tf from Liphistius, returned to subspecies).
H. xianensis Zhu & Wang, 1983: 131, f. 1-7 (Dm).
H. s. Platnick & Sedgwick, 1984: 4 (Tf from Liphistius).
H. s. Song & Haupt, 1984: 447, f. 1c-d, 2b, 2d, 3b (mf, elevated to species, S).
H. xianensis Wang & Zhu, 1984: 403, f. 1-7 (Df).
H. s. Song, Zhu & Chen, 1999: 33, f. 15E, L-M (mf).
Abcathela s. Ono, 2000: 150.
Sinothela s. Haupt, 2003: 71 (Tmf, f. 62E from Heptathela).

f shei Xu & Yin, 2001....................China [urn:lsid:amnh.org:spidersp:037421]
H. s. Xu & Yin, 2001a: 8, f. 1-8 (Df).
H. s. Yin et al., 2012: 117, f. 5a-h, 3-18c-e (f).

mf sinensis Bishop & Crosby, 1932....................China [urn:lsid:amnh.org:spidersp:000023]
H. s. Bishop & Crosby, 1932: 5, f. 1-7 (Df).
H. s. Bristowe, 1933a: 1055, f. 12.
H. s. Wen & Zhu, 1980: 39, f. 1.1-9 (f, Dm).
H. s. Haupt, 1983: 285, f. 9c (f).
H. s. Hu, 1984: 26, f. 24-26 (mf).
H. s. Song & Haupt, 1984: 445, f. 1a-b, 2a, 3a (m).
H. s. Zhu et al., 1985: 48, f. 39a-g (mf).
H. s. Zhang, 1987: 44, f. 30.1-5 (mf).
H. s. Zhao, 1993: 53, f. 19a-c (mf).
H. s. Song, Zhu & Chen, 1999: 33, f. 15F, N-O (mf).
Abcathela s. Ono, 2000: 150.
H. s. Song, Zhu & Chen, 2001: 46, f. 17A-E (mf).
Sinothela s. Haupt, 2003: 71, f. 62F (Tmf from Heptathela).
Songthela s. Zhu & Zhang, 2011: 30, f. 10A-D (mf).

mf tomokunii Ono, 1997....................Vietnam [urn:lsid:amnh.org:spidersp:000025]
H. t. Ono, 1997a: 24, f. 1-8 (Dmf).
H. t. Schwendinger & Ono, 2011: 613, f. 39-43 (mf).

m tonkinensis (Bristowe, 1933)....................Vietnam [urn:lsid:amnh.org:spidersp:000026]
Liphistius birmanicus Simon, 1909e: 70 (m, misidentified).
Liphistius birmanicus Berland, 1932b: 323, f. 442-443 (m, misidentified).
Liphistius t. Bristowe, 1933a: 1022, 1027, f. 8f (Dm).
Liphistius birmanicus Bristowe, 1938b: 661 (m, misidentified).
H. t. Haupt, 1983: 284, f. 8a-b, 8d, 13e (Tm from Liphistius).
H. t. Platnick & Sedgwick, 1984: 3 (Tm from Liphistius).
Vinathela tonkingensis Ono, 2000: 150.
Nanthela t. Haupt, 2003: 69, f. 51A-B, 62C (Tm from Heptathela).
H. t. Schwendinger & Ono, 2011: 611, f. 34-38 (m).

f wosanensis Wang & Jiao, 1995....................China [urn:lsid:amnh.org:spidersp:038358]
H. w. Wang & Jiao, 1995: 80, f. A-D (Df).

f xianningensis Yin et al., 2002....................China [urn:lsid:amnh.org:spidersp:037423]
H. x. Yin et al., 2002: 19, f. 9-16 (Df).

f yaginumai Ono, 1998....................Japan [urn:lsid:amnh.org:spidersp:000027]
H. y. Ono, 1998: 20, f. 19-20 (Df).
H. y. Ono, 2009a: 81, f. 66-67 (f).

mf yakushimaensis Ono, 1998....................Japan [urn:lsid:amnh.org:spidersp:000028]
H. y. Ono, 1998: 23, f. 27-32 (Dmf).
H. kimurai y. Haupt, 2003: 69, f. 54M, 52I (mf).
H. y. Ono, 2009a: 83, f. 85-90 (mf).

mf yanbaruensis Haupt, 1983....................Okinawa [urn:lsid:amnh.org:spidersp:000016]
H. kimurai y. Haupt, 1983: 284, f. 7c-d, 12d, 13d, 13f (Dmf).
H. kimurai y. Haupt, 1984: 166, f. 7 (f).
H. kimurai y. Haupt, 2003: 69, f. 49C-D, 53F-I, K, 54P-U, 55.4-6, 62H (mf).
H. y. Ono, 2009a: 80, f. 53-59 (mf, elevated to species).

f yunnanensis Song & Haupt, 1984....................China [urn:lsid:amnh.org:spidersp:000029]
H. y. Song & Haupt, 1984: 449, f. 3f (Df).
H. y. Song, Zhu & Chen, 1999: 33, f. 15G (f).
Abcathela y. Ono, 2000: 150.