The World Spider Catalog, V15 by N. I. Platnick (c) 2000 - 2014 AMNH

Family: THOMISIDAE

Gen. Heriaeus Simon, 1875 [urn:lsid:amnh.org:spidergen:02680]

N.B.: type species is Thomisus hirtus Latreille, 1819 by ICZN Opinion 1488.

In synonymy:
H. claveatus (Walckenaer, 1837, T from Thomisus, where "nicht zu deuten!" per Roewer) = H. hirtus (Latreille, 1819) (Denis, 1957c: 246).
H. fimbriatus Lawrence, 1942 = H. crassispinus Lawrence, 1942 (Niekerk & Dippenaar-Schoeman, 2013: 459).
H. kumaonensis (Tikader, 1980, T from Misumenops) = H. horridus Tyschchenko, 1965 (Lehtinen, 2005a: 179).
H. propinquus Kulczyn'ski, 1903 = H. simoni Kulczyn'ski, 1903 (Loerbroks, 1983: 107).
H. sareptanus Loerbroks, 1983 = H. horridus Tyschchenko, 1965 (Marusik & Logunov, 1995a: 136).

Nomina dubia:
H. difficilis Strand, 1906b: 626 (f, Tunisia) -- Loerbroks, 1983: 133.
H. melanotrichus Simon, 1903e: 96 (j, Equatorial Guinea) -- Loerbroks, 1983: 133.

mf algericus Loerbroks, 1983....................Algeria [urn:lsid:amnh.org:spidersp:030429]
H. a. Loerbroks, 1983: 123, f. 17, 69-72 (Dmf).

mf allenjonesi Niekerk & Dippenaar-Schoeman, 2013....................South Africa [urn:lsid:amnh.org:spidersp:046592]
H. a. Niekerk & Dippenaar-Schoeman, 2013: 453, f. 2, 14-17 (Dmf).

mf antoni Niekerk & Dippenaar-Schoeman, 2013....................Senegal, Yemen [urn:lsid:amnh.org:spidersp:046593]
H. a. Niekerk & Dippenaar-Schoeman, 2013: 455, f. 3, 18-21 (Dmf).

mf buffoni (Audouin, 1826)....................North Africa, Israel [urn:lsid:amnh.org:spidersp:030430]
Thomisus b. Audouin, 1826: 396, pl. 6, f. 10 (Dm).
Thomisus villosus Lucas, 1846: 192, pl. 10, f. 8 (Df).
H. b. Kulczyn'ski, 1903b: 652, pl. 1, f. 34 (f).
H. b. Levy, 1973: 138, f. 63-66 (mf).
H. buffonii Loerbroks, 1983: 125, f. 21, 73-76 (mf).
H. b. Levy, 1985b: 51, f. 67-70 (mf).

mf buffonopsis Loerbroks, 1983....................Central Asia [urn:lsid:amnh.org:spidersp:030431]
H. b. Loerbroks, 1983: 127, f. 22, 77-80 (Dmf).
H. b. Utochkin, 1985: 109, f. 4, 25-26 (m).

f capillatus Utochkin, 1985....................Kazakhstan [urn:lsid:amnh.org:spidersp:030432]
H. c. Utochkin, 1985: 105, f. 2, 21 (Df).

f charitonovi Utochkin, 1985....................Central Asia [urn:lsid:amnh.org:spidersp:030433]
H. c. Utochkin, 1985: 107, f. 1, 22-23 (Df).

mf concavus Tang & Li, 2010....................China [urn:lsid:amnh.org:spidersp:043677]
H. c. Tang & Li, 2010b: 12, f. 4A-D, 5A-E, 6A-D (Dmf).

mf convexus Tang & Li, 2010....................China [urn:lsid:amnh.org:spidersp:043678]
H. c. Tang & Li, 2010b: 15, f. 7A-D, 8A-E, 9A-D (Dmf).

mf copricola Niekerk & Dippenaar-Schoeman, 2013....................South Africa [urn:lsid:amnh.org:spidersp:046594]
H. c. Niekerk & Dippenaar-Schoeman, 2013: 456, f. 4, 22-25 (Dmf).

mf crassispinus Lawrence, 1942....................Central, East, Southern Africa [urn:lsid:amnh.org:spidersp:030434]
H. c. Lawrence, 1942: 159, f. 13a-b (Df).
H. fimbriatus Lawrence, 1942: 160, f. 14 (Dm) [urn:lsid:amnh.org:spidersp:030437].
H. c. Loerbroks, 1983: 131, f. 93-94 (f).
H. fimbriatus Loerbroks, 1983: 132, f. 90-92 (m).
H. c. Niekerk & Dippenaar-Schoeman, 2013: 459, f. 5, 26-29 (f, Sm).

f delticus Utochkin, 1985....................Russia [urn:lsid:amnh.org:spidersp:030435]
H. d. Utochkin, 1985: 107, f. 3, 24 (Df).

f fedotovi Charitonov, 1946....................Central Asia [urn:lsid:amnh.org:spidersp:030436]
H. f. Charitonov, 1946: 27, f. 46 (Df; see also Charitonov, 1969: 116).
H. f. Utochkin, 1985: 112, f. 8, 20 (f).

mf foordi Niekerk & Dippenaar-Schoeman, 2013....................South Africa [urn:lsid:amnh.org:spidersp:046595]
H. f. Niekerk & Dippenaar-Schoeman, 2013: 461, f. 6, 30-33 (Dmf).

mf graminicola (Doleschall, 1852)....................Europe to Central Asia [urn:lsid:amnh.org:spidersp:030438]
Thomisus g. Doleschall, 1852: 647.
H. savignyi Becker, 1882b: 191, pl. 18, f. 4 (mf, misidentified).
H. hirtus Vilbaste, 1969: 41, f. 36A-B, 37A-B (mf, misidentified).
H. hirtus Tyschchenko, 1971: 114, f. 277 (m, misidentified).
H. g. Loerbroks, 1983: 100, f. 2-5, 15 (mf, removed from S of H. hirtus).
H. g. Utochkin, 1985: 112, f. 7, 27-30 (mf).
H. g. Heimer & Nentwig, 1991: 470, f. 1235 (mf).

mf hirtus (Latreille, 1819) *....................Europe to Georgia [urn:lsid:amnh.org:spidersp:030439]
Thomisus h. Latreille, 1819: 41 (D).
Thomisus hirsutus Walckenaer, 1826: 85 (lapsus).
Thomisus pilosus Risso, 1826: 172 (D).
Thomisus villosus Walckenaer, 1837: 535 (Df).
Pachyptile villosa Simon, 1864: 433.
Misumena hirsuta Thorell, 1873: 539.
Misumena villosa Thorell, 1873: 539.
Misumena hirta Pavesi, 1875: 127.
H. savignyi Simon, 1875a: 205, pl. 7, f. 6 (Dmf).
Misumena savignyi Cantoni, 1881: 286.
H. savignyi Kulczyn'ski, 1903b: 653, pl. 1, f. 24, 32 (mf).
H. h. Simon, 1932: 796, 870, f. 1168-1169 (mf).
H. h. Denis, 1957c: 246 (S).
H. h. Loerbroks, 1983: 103, f. 1, 13, 15, 25-28 (mf).
H. h. Roberts, 1995: 157, f. (mf).
H. h. Mcheidze, 1997: 136, f. 211 (m).
H. h. Roberts, 1998: 165, f. (mf).

mf horridus Tyschchenko, 1965....................Russia, Central Asia, India [urn:lsid:amnh.org:spidersp:030440]
H. h. Tyschchenko, 1965: 698, f. 4 (Dmf).
Misumenops kumaonensis Tikader, 1980a: 157, f. 223-224 (Df).
H. h. Loerbroks, 1983: 131, f. 88-89 (mf).
H. sareptanus Loerbroks, 1983: 128, f. 23, 81-84 (Dmf).
H. sareptanus Marusik & Logunov, 1990: 52, f. 57-58 (m).
H. h. Marusik & Logunov, 1995a: 136, f. 1-3 (mf, S).
H. h. Lehtinen, 2005a: 179 (S).
H. h. Marusik & Kovblyuk, 2011: 253, f. 37.16, 23 (mf).

f latifrons Lessert, 1919....................Tanzania [urn:lsid:amnh.org:spidersp:030441]
H. l. Lessert, 1919: 137, pl. 2, f. 1, 9 (Df).
H. l. Niekerk & Dippenaar-Schoeman, 2013: 463, f. 7, 34 (f).

m madagascar Niekerk & Dippenaar-Schoeman, 2013....................Madagascar [urn:lsid:amnh.org:spidersp:046596]
H. m. Niekerk & Dippenaar-Schoeman, 2013: 464, f. 8, 35-36 (Dm).

mf maurusius Loerbroks, 1983....................Morocco [urn:lsid:amnh.org:spidersp:030442]
H. m. Loerbroks, 1983: 117, f. 51-56 (Dmf).

mf mellotteei Simon, 1886....................China, Korea, Japan [urn:lsid:amnh.org:spidersp:030443]
H. mellottei Simon, 1886b: 177 (Dj; patronym for Mellottée, per Ono, 2011c: 125).
H. melloteei Bösenberg & Strand, 1906: 257, pl. 10, f. 163 (Df).
H. melloteei Saito, 1939: 84, f. 10(5) (f).
H. melloteei Yaginuma, 1960: 99, f. 85.1 (m).
H. melloteei Yaginuma, 1971: 99, f. 85.1 (m).
H. melloteei Hu & Guo, 1982: 137, f. 1-6 (mf).
H. oblongus Shi & Zhu, 1982: 64, f. 2a-e (m, misidentified).
H. melloteei Loerbroks, 1983: 114, f. 14, 47-50 (mf, S H. oblongus, rejected by Ono, 1988c: 172).
H. melloteei Hu, 1984: 316, f. 333.1-6 (mf).
H. oblongus Hu, 1984: 316, f. 334.1-5 (m, misidentified).
H. oblongus Zhu et al., 1985: 170, f. 151a-e (mf, misidentified).
H. melloteei Yaginuma, 1986a: 208, f. 115.2 (mf).
H. oblongus Song, 1987: 254, f. 209 (mf, misidentified).
H. mellottei Zhang, 1987: 204, f. 176.1-4 (mf).
H. oblongus Li, 1987: 223, f. 1-4 (m, misidentified).
H. setiger Hu & Li, 1987b: 312, f. 38.1-5 (mf, misidentified per Song & Zhu, 1997: 154).
H. mellottei Ono, 1988c: 172, f. 175-179 (mf).
H. mellottei Chikuni, 1989b: 139, f. 8 (mf).
H. m. Chen & Gao, 1990: 168, f. 213a-b (mf).
H. mellottei Zhao, 1993: 359, f. 178a-c (mf).
H. melloteei Song & Zhu, 1997: 154, f. 109A-D (mf).
H. melloteei Song, Zhu & Chen, 1999: 481, f. 274J, 277F, O (mf).
H. mellottei Hu, 2001: 339, f. 204.1-5 (mf).
H. melloteei Kim & Gwon, 2001: 25, f. 19-22 (mf).
H. m. Song, Zhu & Chen, 2001: 390, f. 257A-D (mf).
H. melloteei Namkung, 2002: 522, f. 42.5a-b (mf).
H. melloteei Namkung, 2003: 525, f. 42.5a-b (mf).
H. m. Ono, 2009a: 525, f. 283-286 (mf).
H. melloteei Zhu & Zhang, 2011: 443, f. 317A-D (mf).

m muizenberg Niekerk & Dippenaar-Schoeman, 2013....................South Africa [urn:lsid:amnh.org:spidersp:046597]
H. m. Niekerk & Dippenaar-Schoeman, 2013: 465, f. 9, 37-38 (Dm).

mf numidicus Loerbroks, 1983....................Morocco, Algeria [urn:lsid:amnh.org:spidersp:030444]
H. setiger Simon, 1875a: 208, pl. 7, f. 3-4 (mf, misidentified).
H. setiger Kulczyn'ski, 1903b: 653, pl. 1, f. 21, 30 (mf, misidentified).
H. setiger Simon, 1932: 796, 871, f. 1170-1171 (mf, misidentified).
H. n. Loerbroks, 1983: 113, f. 17, 43-46 (Dmf).

mf oblongus Simon, 1918....................Palearctic [urn:lsid:amnh.org:spidersp:043808]
H. hirsutus Simon, 1875a: 206, pl. 7, f. 5 (mf, misidentified).
H. hirsutus Chyzer & Kulczyn'ski, 1891: 85, pl. 3, f. 16 (mf, misidentified).
H. hirsutus Bösenberg, 1902: 368, pl. 34, f. 542 (mf, misidentified).
H. hirsutus Kulczyn'ski, 1903b: 653, pl. 1, f. 22-23, 31 (mf, misidentified).
H. hirsutus Lessert, 1910b: 355, f. 181 (mf, misidentified).
H. o. Simon, 1918a: 51 (Dmf).
H. o. Simon, 1932: 796, 871, f. 1166-1167 (mf).
H. o. Azheganova, 1968: 117, f. 254, 297 (mf).
H. o. Tyschchenko, 1971: 114, f. 274 (m).
H. o. Miller, 1971: 114, pl. XIII, f. 24 (m).
H. o. Barrientos & Urones, 1985: 361, f. 7a-b (f).
H. melloteei Utochkin, 1985: 109, f. 10, 17-19 (mf, misidentified).
H. mellottei Heimer & Nentwig, 1991: 470, f. 1236 (mf, misidentified).
H. mellottei Danilov, 1993a: 61, f. 4-5 (mf, misidentified).
H. mellottei Roberts, 1998: 166, f. (mf, misidentified).

mf orientalis Simon, 1918....................Greece, Turkey, Ukraine [urn:lsid:amnh.org:spidersp:030445]
Thomisus hirtus C. L. Koch, 1837a: 42, f. 275-276 (Dmf, preoccupied).
H. hirtus Kulczyn'ski, 1903b: 653, pl. 1, f. 19-20, 28, 35 (mf, misidentified).
H. o. Simon, 1918a: 51 (replacement name).
H. o. Loerbroks, 1983: 120, f. 19, 61-64 (mf).
H. o. Utochkin, 1985: 109, f. 5-6, 11-13 (mf).
H. melloteei Roberts, 1995: 157, f. (mf, misidentified).
H. oblongus Mcheidze, 1997: 137, f. 212-214 (mf).

mf peterwebbi Niekerk & Dippenaar-Schoeman, 2013....................Namibia, South Africa [urn:lsid:amnh.org:spidersp:046598]
H. transvaalicus Loerbroks, 1983: 130, f. 24, 85-87 (m, misidentified).
H. p. Niekerk & Dippenaar-Schoeman, 2013: 466, f. 10, 39-42 (Dmf).

mf pilosus Nosek, 1905....................Turkey [urn:lsid:amnh.org:spidersp:030446]
H. p. Nosek, 1905b: 135 (Dmf).
H. p. Loerbroks, 1983: 119, f. 18, 57-60 (mf).

mf setiger (O. P.-Cambridge, 1872)....................Palearctic [urn:lsid:amnh.org:spidersp:030447]
Thomisus s. O. P.-Cambridge, 1872a: 307, pl. 14, f. 15 (Dm).
H. s. Tyschchenko, 1971: 114, f. 276 (m).
H. s. Levy, 1973: 136, f. 59-62 (m, Df).
H. s. Loerbroks, 1983: 110, f. 16, 37-42 (mf).
H. s. Levy, 1985b: 49, f. 63-66 (mf).

mf simoni Kulczyn'ski, 1903....................Palearctic [urn:lsid:amnh.org:spidersp:030448]
H. simonii Kulczyn'ski, 1903b: 654, pl. 1, f. 25, 29 (Dmf).
H. propinquus Kulczyn'ski, 1903b: 657, pl. 1, f. 27 (Df).
H. simonii Loerbroks, 1983: 107, f. 15, 29-36 (mf, S).
H. somonii Qiu, 1983: 93, f. 13.4a-c (mf).
H. simonii Utochkin, 1985: 112, f. 7-10, 14-16 (mf).

f sossusvlei Niekerk & Dippenaar-Schoeman, 2013....................Namibia, South Africa [urn:lsid:amnh.org:spidersp:046599]
H. s. Niekerk & Dippenaar-Schoeman, 2013: 468, f. 43-44 (Df).

mf spinipalpus Loerbroks, 1983....................Eastern Mediterranean [urn:lsid:amnh.org:spidersp:030449]
H. s. Loerbroks, 1983: 122, f. 20, 65-68 (Dmf).
H. s. Ono & Martens, 2005: 121, f. 71-74 (f).

mf transvaalicus Simon, 1895....................South Africa [urn:lsid:amnh.org:spidersp:030450]
H. t. Simon, 1895b: 438 (Df).
H. t. Loerbroks, 1983: 130, f. 24, 85-87 (f, not m, =H. peterwebbi).
H. t. Niekerk & Dippenaar-Schoeman, 2013: 469, f. 1, 11, 45-48 (f, Dm).

mf xanderi Niekerk & Dippenaar-Schoeman, 2013....................Tanzania, South Africa [urn:lsid:amnh.org:spidersp:046599]
H. x. Niekerk & Dippenaar-Schoeman, 2013: 471, f. 12, 49-52 (Dmf).

mf zanii Niekerk & Dippenaar-Schoeman, 2013....................South Africa [urn:lsid:amnh.org:spidersp:046600]
H. z. Niekerk & Dippenaar-Schoeman, 2013: 472, f. 13, 53-56 (Dmf).