The World Spider Catalog, V15 by N. I. Platnick (c) 2000 - 2014 AMNH

Family: GNAPHOSIDAE

Gen. Herpyllus Hentz, 1832 [urn:lsid:amnh.org:spidergen:02432]

N.B.: considered a senior synonym of Bonna O. P.-Cambridge, 1898 by Platnick & Shadab, 1977b: 4; Old World species probably misplaced.

Transferred to other genera:
H. anatolicus Kamura, 1989 -- see Sanitubius.
H. angustus Banks, 1904 -- see Sergiolus.
H. atopophysis Chamberlin, 1928 -- see Nodocion.
H. cultior (Kulczyn'ski, 1899) -- see Scotophaeus.
H. floridanus (Banks, 1896) -- see Nodocion.
H. haplodrassoides Denis, 1955 -- see Zelotes.
H. irvingi Mello-Leitão, 1944 -- see Cesonia.
H. josephus (Chamberlin & Gertsch, 1940) -- see Cesonia.
H. longipes (Nicolet, 1849) -- see Camillina.
H. pius Chamberlin, 1920 -- see Scotophaeus.
H. striatus (Kishida, 1932) -- see Drassodes.
H. suavis Simon, 1893 -- see Apopyllus.

In synonymy:
H. bryantae (Roewer, 1951, T from Zelotes) = H. ecclesiasticus Hentz, 1832 (Ubick & Roth, 1973a: 5).
H. bryophilus Chamberlin, 1936 = H. emertoni Bryant, 1935 (Platnick & Shadab, 1977b: 21).
H. cepeus Chamberlin, 1936 = H. cockerelli (Banks, 1901) (Platnick & Shadab, 1977b: 16).
H. cratus Chamberlin, 1922 = H. ecclesiasticus Hentz, 1832 (Platnick & Shadab, 1977b: 7).
H. faxoni Bryant, 1936 = H. emertoni Bryant, 1935 (Platnick & Shadab, 1977b: 21).
H. itamus Chamberlin, 1936 = H. emertoni Bryant, 1935 (Platnick & Shadab, 1977b: 21).
H. lesserti (Schenkel, 1950, T from Poecilochroa) = H. hesperolus Chamberlin, 1928 (Platnick & Shadab, 1981e: 7).
H. piedicus Chamberlin & Woodbury, 1929 = H. propinquus (Keyserling, 1887) (Platnick & Shadab, 1977b: 9).

Homonym replaced:
H. validus (Banks, 1896) -- see H. hesperolus.

Nomina dubia:
H. pygmaeus Hentz, 1847: 459 (USA) -- Ubick & Roth, 1973b: 5.
H. ramulosus Hentz, 1847: 459 (USA) -- Ubick & Roth, 1973b: 5.
H. vespa Hentz, 1847: 458, pl. 24, f. 13 (f, USA) -- Platnick & Shadab, 1977b: 6.

Nomen nudum:
H. vasifer (Walckenaer, 1805: 46) -- Platnick & Shadab, 1977b: 4.

mf australis (Holmberg, 1881)....................Argentina [urn:lsid:amnh.org:spidersp:027391]
Drassus a. Holmberg, 1881b: 136, pl. 3, f. 4 (Dm).
H. a. Mello-Leitão, 1940b: 46, f. 44-45 (m, Df).

mf bensonae Fox, 1938....................Mexico [urn:lsid:amnh.org:spidersp:027392]
H. b. Fox, 1938a: 232, pl. 1, f. 7 (Dm).
H. b. Platnick & Shadab, 1977b: 26, f. 71-74 (m, Df).

f brachet Platnick & Shadab, 1977....................Mexico [urn:lsid:amnh.org:spidersp:027393]
H. b. Platnick & Shadab, 1977b: 37, f. 113-114 (Df).

mf bubulcus Chamberlin, 1922....................USA, Mexico [urn:lsid:amnh.org:spidersp:027394]
H. b. Chamberlin, 1922: 150 (Df).
H. b. Platnick & Shadab, 1977b: 29, f. 85-90 (f, Dm).

f calcuttaensis Biswas, 1984....................India [urn:lsid:amnh.org:spidersp:027395]
H. c. Biswas, 1984a: 123, f. 7-10 (Df).

mf coahuilanus Gertsch & Davis, 1940....................Mexico [urn:lsid:amnh.org:spidersp:027396]
H. c. Gertsch & Davis, 1940b: 5, f. 1-2 (Dmf).
H. c. Platnick & Shadab, 1977b: 34, f. 99-102 (mf).

mf cockerelli (Banks, 1901)....................USA, Mexico [urn:lsid:amnh.org:spidersp:027397]
Prosthesima c. Banks, 1901b: 571, pl. 33, f. 2 (Dmf).
H. c. Banks, 1910: 8.
H. cepeus Chamberlin, 1936b: 2, f. 13 (Df).
H. c. Platnick & Shadab, 1977b: 15, f. 31-36 (mf, S).

mf convallis Chamberlin, 1936....................USA, Mexico [urn:lsid:amnh.org:spidersp:027398]
H. c. Chamberlin, 1936a: 2, f. 1-3 (Dmf).
H. c. Platnick & Shadab, 1977b: 27, f. 69-70, 75-78 (mf).

m coreanus Paik, 1992....................Korea [urn:lsid:amnh.org:spidersp:027399]
H. c. Paik, 1992b: 133, f. 1-10 (Dm).

mf ecclesiasticus Hentz, 1832 *....................North America [urn:lsid:amnh.org:spidersp:027400]
H. e. Hentz, 1832: 102 (Df).
Drassus vasifer Walckenaer, 1837: 620 (Df).
H. e. Hentz, 1847: 455, pl. 24, f. 2 (f).
Prosthesima bimaculata Keyserling, 1887b: 433, pl. 6, f. 9 (Dm).
Prosthesima ecclesiastica Emerton, 1890: 173, pl. 3, f. 7 (f, Dm).
Prosthesima ecclesiastica Emerton, 1902: 5, f. 17-18 (f).
Melanophora bimaculata Bryant, 1908: 7.
H. vasifer Comstock, 1912: 318, f. 308.
H. cratus Chamberlin, 1922: 150 (Df).
H. vasifer Kaston, 1948: 349, f. 1165-1169 (mf).
Zelotes bryantae Roewer, 1951: 444 (replacement name for Prosthesima bimaculata).
H. e. Ubick & Roth, 1973a: 5 (removed mf from S of H. vasifer, S).
H. e. Platnick & Shadab, 1977b: 7, f. 1-8 (mf, S).
H. e. Heiss & Allen, 1986: 60, f. 99-101 (mf).
H. e. Platnick & Dondale, 1992: 273, f. 14, 433-436 (mf).
H. e. Paquin & Dupérré, 2003: 79, f. 685-687 (mf).
H. e. Murphy, 2007: 46, f. 278-279 (mf).

mf emertoni Bryant, 1935....................USA [urn:lsid:amnh.org:spidersp:027401]
H. e. Bryant, 1935a: 73, pl. 5, f. 1-2 (Df).
H. faxoni Bryant, 1936a: 96, pl. 3, f. 6 (Df).
H. e. Chamberlin, 1936b: 1, f. 8 (Dm).
H. bryophilus Chamberlin, 1936b: 1, f. 9 (Df).
H. itamus Chamberlin, 1936b: 2, f. 10 (Df).
H. e. Platnick & Shadab, 1977b: 21, f. 51-56 (mf, S).

mf excelsus Fox, 1938....................USA, Mexico [urn:lsid:amnh.org:spidersp:027402]
H. e. Fox, 1938a: 232, pl. 2, f. 8 (Df).
H. e. Platnick & Shadab, 1977b: 12, f. 15-20 (f, Dm).

f fidelis (O. P.-Cambridge, 1898)....................Mexico [urn:lsid:amnh.org:spidersp:027403]
Bonna f. O. P.-Cambridge, 1898a: 250, pl. 33, f. 1 (Df).
H. f. Platnick & Shadab, 1977b: 37, f. 115-116 (f).

mf frio Platnick & Shadab, 1977....................Mexico [urn:lsid:amnh.org:spidersp:027404]
H. f. Platnick & Shadab, 1977b: 14, f. 27-30 (Dmf).

mf gertschi Platnick & Shadab, 1977....................USA, Mexico [urn:lsid:amnh.org:spidersp:027405]
H. g. Platnick & Shadab, 1977b: 35, f. 57-58, 103-106 (Dmf).

f giganteus Platnick & Shadab, 1977....................Mexico [urn:lsid:amnh.org:spidersp:027406]
H. g. Platnick & Shadab, 1977b: 38, f. 117-118 (Df).

f goaensis Tikader, 1982....................India [urn:lsid:amnh.org:spidersp:027407]
H. g. Tikader, 1982b: 460, f. 372-376 (Df).
H. g. Gajbe, 2007: 478, f. 162-166 (f).

mf hesperolus Chamberlin, 1928....................North America [urn:lsid:amnh.org:spidersp:027408]
Prosthesima valida Banks, 1896e: 62 (Dmf, preoccupied).
H. validus Banks, 1904b: 337, pl. 39, f. 25, 29 (mf).
H. h. Chamberlin, in Chamberlin & Gertsch, 1928: 176 (replacement name).
Sergiolus lesserti Schenkel, 1950b: 42, f. 9 (Df).
Poecilochroa lesserti Ubick & Roth, 1973a: 7 (Tf from Sergiolus).
H. h. Platnick & Shadab, 1977b: 23, f. 63-68 (mf).
H. h. Platnick & Shadab, 1981e: 7 (S).
H. h. Platnick & Dondale, 1992: 271, f. 429-432 (mf).

mf iguala Platnick & Shadab, 1977....................Mexico [urn:lsid:amnh.org:spidersp:027409]
H. i. Platnick & Shadab, 1977b: 23, f. 59-62 (Dmf).

f lativulvus Denis, 1958....................Afghanistan [urn:lsid:amnh.org:spidersp:027410]
H. lativulva Denis, 1958b: 88, f. 6 (Df).

f malkini Platnick & Shadab, 1977....................Mexico [urn:lsid:amnh.org:spidersp:027411]
H. m. Platnick & Shadab, 1977b: 37, f. 111-112 (Df).

f paropanisadensis Denis, 1958....................Afghanistan [urn:lsid:amnh.org:spidersp:027412]
H. p. Denis, 1958b: 86, f. 3 (Df).

mf perditus (Banks, 1898)....................Mexico [urn:lsid:amnh.org:spidersp:027413]
Drassodes p. Banks, 1898b: 215, pl. 13, f. 7 (Dmf).
H. p. Platnick & Shadab, 1976a: 4 (Tmf from Drassodes).
H. p. Platnick & Shadab, 1977b: 32, f. 95-98 (mf).

mf perote Platnick & Shadab, 1977....................Mexico [urn:lsid:amnh.org:spidersp:027414]
H. p. Platnick & Shadab, 1977b: 35, f. 107-110 (Dmf).

mf pictus (F. O. P.-Cambridge, 1899)....................Mexico [urn:lsid:amnh.org:spidersp:027415]
Scotophaeus p. F. O. P.-Cambridge, 1899a: 59, pl. 4, f. 14 (Df).
H. p. Ubick & Roth, 1973c: 2 (Tf from Scotophaeus).
H. p. Platnick & Shadab, 1977b: 19, f. 47-50 (f, Dm).

mf propinquus (Keyserling, 1887)....................North America [urn:lsid:amnh.org:spidersp:027416]
Prosthesima propinqua Keyserling, 1887b: 430, pl. 6, f. 7 (Df).
H. californicus Banks, 1904a: 110, f. 11 (Df).
H. piedicus Chamberlin & Woodbury, 1929: 132, pl. 2, f. 6-8 (Dm).
H. p. Chamberlin, 1936a: 1.
H. p. Platnick & Shadab, 1977b: 9, f. 9-14, 130 (f, Sm).
H. p. Platnick & Dondale, 1992: 275, f. 437-440 (mf).
H. p. Zakharov & Ovtcharenko, 2013: 16, f. 2a-b (m).

f proximus Denis, 1958....................Turkmenistan, Afghanistan [urn:lsid:amnh.org:spidersp:027417]
H. p. Denis, 1958b: 88, f. 5 (Df).

mf regnans Chamberlin, 1936....................USA [urn:lsid:amnh.org:spidersp:027418]
H. r. Chamberlin, 1936b: 2, f. 14 (Df).
H. r. Platnick & Shadab, 1977b: 31, f. 91-94 (f, Dm).

mf reservatus Chamberlin, 1936....................USA, Mexico [urn:lsid:amnh.org:spidersp:027419]
H. r. Chamberlin, 1936b: 3, f. 15 (Df).
H. r. Platnick & Shadab, 1977b: 19, f. 21-22, 43-46 (f, Dm).

mf scholasticus Chamberlin, 1922....................USA [urn:lsid:amnh.org:spidersp:027420]
H. s. Chamberlin, 1922: 149 (Dm).
H. s. Platnick & Shadab, 1977b: 18, f. 37-42 (mf).

mf schwarzi (Banks, 1901)....................USA [urn:lsid:amnh.org:spidersp:027421]
Prosthesima s. Banks, 1901a: 582, pl. 22, f. 7 (Df).
H. s. Banks, 1910: 8.
H. s. Platnick & Shadab, 1977b: 28, f. 79-84 (f, Dm).

mf sherus Platnick & Shadab, 1977....................Mexico [urn:lsid:amnh.org:spidersp:027422]
H. s. Platnick & Shadab, 1977b: 13, f. 23-26 (Dmf).

f vicinus Denis, 1958....................Afghanistan [urn:lsid:amnh.org:spidersp:027423]
H. v. Denis, 1958b: 87, f. 4 (Df).