The World Spider Catalog, V15 by N. I. Platnick (c) 2000 - 2014 AMNH

Family: THOMISIDAE

Gen. Lysiteles Simon, 1895 [urn:lsid:amnh.org:spidergen:02693]

Transferred to other genera:
L. guangxiensis He & Hu, 1999 -- see Alcimochthes.

In synonymy:
L. nigrifrons (Saito, 1934) = L. coronatus (Grube, 1861) (Ono, 1988c: 133).
L. sapporensis (Saito, 1934) = L. coronatus (Grube, 1861) (Ono, 1988c: 133).
L. takashimai (Uyemura, 1937) = L. coronatus (Grube, 1861) (Ono, 1985a: 23).
L. xianensis Song & Wang, 1991 = L. minimus (Schenkel, 1953) (Song, 1995: 125).

mf ambrosii Ono, 2001....................Bhutan, China [urn:lsid:amnh.org:spidersp:038182]
L. a. Ono, 2001b: 217, f. 28-33 (Dmf).
L. a. Tang et al., 2007: 64, f. 41-48 (mf).

f amoenus Ono, 1980....................Bhutan, Taiwan [urn:lsid:amnh.org:spidersp:030482]
L. a. Ono, 1980a: 214, f. 31-34 (Df).
L. a. Song & Zhu, 1997: 120, f. 83A-D (f).
L. a. Song, Zhu & Chen, 1999: 481, f. 277G (f).
L. a. Ono, 2001b: 212, f. 14-15 (f).

mf anchorus Zhu, Lian & Ono, 2004....................China [urn:lsid:amnh.org:spidersp:040717]
L. a. Zhu, Lian & Ono, 2004: 53, f. 1-7 (Dmf).

m annapurnus Ono, 1979....................Nepal [urn:lsid:amnh.org:spidersp:030483]
L. a. Ono, 1979: 105, f. 35-37 (Dm).

f arcuatus Tang et al., 2008....................China [urn:lsid:amnh.org:spidersp:041566]
L. a. Tang et al., 2008b: 5, f. 1a-e (Df).

f auriculatus Tang et al., 2008....................China [urn:lsid:amnh.org:spidersp:041567]
L. a. Tang et al., 2008b: 6, f. 2a-e (Df).

mf badongensis Song & Chai, 1990....................China [urn:lsid:amnh.org:spidersp:030484]
L. b. Song & Chai, 1990: 367, f. 4A-D (Dmf).
L. b. Song & Li, 1997: 422, f. 27A-D (mf).
L. b. Song & Zhu, 1997: 122, f. 85A-D (mf).
L. b. Song, Chen & Zhu, 1997: 1730, f. 37a-d (mf).
L. b. Song, Zhu & Chen, 1999: 481, f. 278A, I (mf).
L. b. Zhu & Zhang, 2011: 445, f. 318A-D (mf).

mf bhutanus Ono, 2001....................Bhutan, China [urn:lsid:amnh.org:spidersp:038183]
L. b. Ono, 2001b: 222, f. 44-51 (Dmf).
L. b. Tang et al., 2007: 66, f. 49-58 (mf).

m boteus Barrion & Litsinger, 1995....................Philippines [urn:lsid:amnh.org:spidersp:030485]
L. b. Barrion & Litsinger, 1995: 262, f. 153a-f (Dm).

f brunettii (Tikader, 1962)....................India [urn:lsid:amnh.org:spidersp:030486]
Synaema b. Tikader, 1962b: 578, f. 10 (Df).
Synaema b. Tikader, 1971a: 54, f. 15N-P (f).
L. b. Ono, 1979: 106 (Tf from Synema).
Synaema b. Tikader, 1980a: 134, f. 190-191 (f).

f catulus Simon, 1895 *....................India [urn:lsid:amnh.org:spidersp:030487]
L. c. Simon, 1895b: 434 (Df).
L. c. Ono, 1979: 95, f. 4-7 (f).

mf clavellatus Tang et al., 2008....................China [urn:lsid:amnh.org:spidersp:041568]
L. c. Tang et al., 2008b: 7, f. 3a-l (Dmf).

mf conflatus Tang et al., 2008....................China [urn:lsid:amnh.org:spidersp:041569]
L. c. Tang et al., 2008b: 9, f. 4a-l (Dmf).

mf conicus Tang et al., 2007....................China [urn:lsid:amnh.org:spidersp:040718]
L. c. Tang et al., 2007: 60, f. 8-27 (Dmf).

mf coronatus (Grube, 1861)....................Russia, China, Korea, Japan [urn:lsid:amnh.org:spidersp:030488]
Thomisus c. Grube, 1861: 173 (Df).
Xysticus c. Reimoser, 1919: 195.
Ozyptila nigrifrons Saito, 1934b: 279, pl. 12, f. 7, pl. 14, f. 40 (Dm).
Xysticus sapporensis Saito, 1934b: 280, pl. 12, f. 8, pl. 14, f. 41 (Df).
Oxyptila takashimai Uyemura, 1937c: 153, f. 2a-e, pl. 10 (Df).
Diaea takashimai Yaginuma, 1955c: 15 (Tmf from Ozyptila).
Oxyptila nigrifrons Saito, 1959: 128, f. 161a-d (m).
Diaea takashimai Namkung, 1964: 42, f. 37 (f).
Synaema takashimai Yaginuma, 1967b: 88, f. 1d (Tmf from Diaea).
Synaema takashimai Arita, 1970: 27, f. 11 (f).
Synaema takashimai Yaginuma, 1971: 98, f. 85.4-5 (f).
Synaema takashimai Kobayashi, 1974: 46, f. 4 (f).
L. nigrifrons Ono, 1979: 106 (Tm from Ozyptila).
L. sapporensis Ono, 1979: 106 (Tf from Xysticus).
L. takashimai Ono, 1979: 106 (Tmf from Synema).
L. takashimai Ono, 1980a: 204, f. 1-6 (mf).
L. c. Ono, 1985a: 23, f. 13-15 (Tf from Xysticus, Sm).
Xysticus sapporensis Zhu et al., 1985: 196, f. 178a-c (f).
L. c. Yaginuma, 1986a: 212, f. 118.1 (mf).
L. c. Ono, 1988c: 133 (S).
L. c. Chikuni, 1989b: 141, f. 14 (mf).
L. c. Ono, Marusik & Logunov, 1990: 14, f. 34-36 (mf).
L. c. Logunov, 1990: 39, f. 4 (m).
L. c. Kim & Gwon, 2001: 26, f. 23-25 (f).
L. c. Namkung, 2002: 525, f. 42.8a-b (mf).
L. c. Namkung, 2003: 528, f. 42.8a-b (mf).
L. c. Ono, 2009a: 520, f. 237-242 (mf).

f corrugus Tang et al., 2008....................China [urn:lsid:amnh.org:spidersp:041570]
L. c.Tang et al., 2008b: 12, f. 5a-g (Df).

f curvatus Tang et al., 2008....................China [urn:lsid:amnh.org:spidersp:041571]
L. c. Tang et al., 2008b: 13, f. 6a-g (Df).

f davidi Tang et al., 2007....................China [urn:lsid:amnh.org:spidersp:040719]
L. d. Tang et al., 2007: 58, f. 1-7 (Df).

mf dentatus Tang et al., 2007....................China [urn:lsid:amnh.org:spidersp:040720]
L. d. Tang et al., 2007: 62, f. 28-40 (Dmf).

mf dianicus Song & Zhao, 1994....................China [urn:lsid:amnh.org:spidersp:030489]
L. d. Song & Zhao, 1994: 114, f. 2A-C (Df).
L. d. Song & Zhu, 1997: 121, f. 84A-C (f).
L. d. Song, Zhu & Chen, 1999: 481, f. 277H (f).
L. d. Tang et al., 2008b: 14, f. 7a-k (f, Dm).

m digitatus Zhang, Zhu & Tso, 2006....................Taiwan [urn:lsid:amnh.org:spidersp:040721]
L. d. Zhang, Zhu & Tso, 2006b: 78, f. 1-3 (Dm).

m distortus Tang et al., 2008....................China [urn:lsid:amnh.org:spidersp:041572]
L. d. Tang et al., 2008b: 17, f. 8a-e (Dm).

mf excultus (O. P.-Cambridge, 1885)....................India, Pakistan [urn:lsid:amnh.org:spidersp:030490]
Synema exculta O. P.-Cambridge, 1885b: 60 (Dmf).
L. e. Marusik, 1993b: 464, f. 19-24 (Tf from Synema, Dm).

mf furcatus Tang & Li, 2010....................China [urn:lsid:amnh.org:spidersp:043018]
L. f. Tang & Li, 2010a: 31, f. 23A-D, 24A-E, 25A-D (Dmf).

f guoi Tang et al., 2008....................China [urn:lsid:amnh.org:spidersp:041573]
L. g. Tang et al., 2008b: 18, f. 9a-e (Df).

mf himalayensis Ono, 1979....................Nepal, Bhutan, China [urn:lsid:amnh.org:spidersp:030491]
L. h. Ono, 1979: 102, f. 29-31 (Dm).
L. h. Ono, 2001b: 225, f. 52-55 (m, Df).
L. h. Tang et al., 2008b: 20, f. 10a-e (f).

f hongkong Song, Zhu & Wu, 1997....................China [urn:lsid:amnh.org:spidersp:030492]
L. h. Song, Zhu & Wu, 1997: 82, f. 3A-B (Df).
L. h. Song, Zhu & Chen, 1999: 481, f. 278B (f).

mf inflatus Song & Chai, 1990....................China [urn:lsid:amnh.org:spidersp:030493]
L. i. Song & Chai, 1990: 368, f. 5A-E (Dmf).
L. i. Song & Li, 1997: 423, f. 28A-E (mf).
L. i. Song & Zhu, 1997: 123, f. 86A-E (mf).
L. i. Song, Zhu & Chen, 1999: 481, f. 274L, 278C, J (mf).
L. i. Yin et al., 2012: 1285, f. 690a-c (f).

mf kunmingensis Song & Zhao, 1994....................Bhutan, China [urn:lsid:amnh.org:spidersp:030494]
L. k. Song & Zhao, 1994: 114, f. 3A-C (Dm).
L. k. Song & Zhu, 1997: 124, f. 87A-C (m).
L. k. Song, Zhu & Chen, 1999: 481, f. 278K (m).
L. k. Ono, 2001b: 214, f. 21-27 (m, Df).
L. k. Tang et al., 2008b: 21, f. 11a-i (mf).

f leptosiphus Tang & Li, 2010....................China [urn:lsid:amnh.org:spidersp:043019]
L. l. Tang & Li, 2010a: 35, f. 26A-F, 27A-B (Df).

f lepusculus Ono, 1979....................Nepal [urn:lsid:amnh.org:spidersp:030495]
L. l. Ono, 1979: 98, f. 14-16 (Df).

f linzhiensis Hu, 2001....................China [urn:lsid:amnh.org:spidersp:038184]
L. l. Hu, 2001: 343, f. 206.1-3 (Df).

m magkalapitus Barrion & Litsinger, 1995....................Philippines [urn:lsid:amnh.org:spidersp:030496]
L. m. Barrion & Litsinger, 1995: 264, f. 154a-g (Dm).

mf maior Ono, 1979....................Russia, Nepal to Japan [urn:lsid:amnh.org:spidersp:030497]
L. maius Ono, 1979: 103, f. 1, 32-34 (Dm).
L. maius Ono, 1980a: 207, f. 7-18 (m, Df).
L. m. Brignoli, 1983c: 609.
L. maius Yaginuma, 1986a: 213, f. 118.2 (mf).
L. maius Ono, 1988c: 137 (contends specific name was a noun in apposition, but the word is a neuter comparative adjective, per H. D. Cameron).
L. maius Chikuni, 1989b: 141, f. 16 (mf).
L. maius Feng, 1990: 178, f. 153.1-2 (f).
L. maius Ono, Marusik & Logunov, 1990: 14, f. 37-41 (mf).
L. maius Danilov, 1993a: 61, f. 6-8 (mf).
L. maius Song & Zhu, 1997: 125, f. 88A-D (mf).
L. maius Song, Zhu & Chen, 1999: 482, f. 278D, L (mf).
L. maius Namkung, 2002: 526, f. 42.9a-b (mf).
L. maius Namkung, 2003: 529, f. 42.9a-b (mf).
L. maius Tang et al., 2008b: 23, f. 12a-l (mf).
L. maius Ono, 2009a: 520, f. 243-248 (mf).

f mandali (Tikader, 1966)....................India, China [urn:lsid:amnh.org:spidersp:030498]
Xysticus m. Tikader, 1966g: 57, f. 4a-b (Df).
Xysticus m. Tikader, 1968b: 110, f. 50-51 (f).
Xysticus m. Tikader, 1971a: 49, f. 14H-J (f).
L. m. Ono, 1979: 106 (Tf from Xysticus).
Xysticus m. Tikader, 1980a: 118, f. 163-164 (f).

mf miniatus Ono, 1980....................Ryukyu Is. [urn:lsid:amnh.org:spidersp:030499]
L. m. Ono, 1980a: 210, f. 22-24 (Df).
L. m. Ono, 1988c: 142, f. 145-147 (Dm).
L. m. Ono, 2009a: 521, f. 254-258 (mf).

mf minimus (Schenkel, 1953)....................China [urn:lsid:amnh.org:spidersp:030500]
Xysticus m. Schenkel, 1953b: 55, f. 27a-b (Dm).
L. m. Ono, 1979: 106 (Tm from Xysticus).
L. xianensis Song & Wang, 1991: 2, f. 2A-C (Dm).
L. m. Song, 1995: 125 (S).
L. m. Song & Zhu, 1997: 126, f. 89A-B (m).
L. m. Song, Zhu & Chen, 1999: 482, f. 278M (m).
L. m. Hu & Zhang, 2011b: 65, f. 1-7 (m, Df).
L. m. Zhu & Zhang, 2011: 445, f. 319A-B (m).

mf minusculus Song & Chai, 1990....................Bhutan, China [urn:lsid:amnh.org:spidersp:030501]
L. m. Song & Chai, 1990: 369, f. 6A-C (Dm).
L. m. Song & Li, 1997: 424, f. 29A-C (m).
L. m. Song & Zhu, 1997: 127, f. 90A-C (m).
L. m. Song, Zhu & Chen, 1999: 482, f. 278N (m).
L. m. Ono, 2001b: 214, f. 16-20 (m).
L. m. Tang & Li, 2010a: 37, f. 28A-D, 29A-C, 30A-D (m, Df).

f montivagus Ono, 1979....................Nepal [urn:lsid:amnh.org:spidersp:030502]
L. m. Ono, 1979: 101, f. 21-24 (Df).

mf niger Ono, 1979....................Nepal, Bhutan, China [urn:lsid:amnh.org:spidersp:030503]
L. n. Ono, 1979: 102, f. 25-28 (Df).
L. n. Ono, 2001b: 219, f. 34-35 (f, Dm).
L. n. Tang et al., 2008b: 26, f. 13a-k (mf).

mf okumae Ono, 1980....................Japan [urn:lsid:amnh.org:spidersp:030504]
L. okumai Ono, 1980a: 209, f. 19-21 (Df).
L. o. Ono, 1988c: 140, f. 142-144 (Dm).
L. o. Chikuni, 1989b: 141, f. 15 (mf).
L. o. Ono, 2009a: 521, f. 249-253 (mf).

mf parvulus Ono, 1979....................Nepal [urn:lsid:amnh.org:spidersp:030505]
L. p. Ono, 1979: 96, f. 8-13 (Dmf).

mf punctiger Ono, 2001....................Bhutan, China [urn:lsid:amnh.org:spidersp:038185]
L. p. Ono, 2001b: 228, f. 62-64 (Df).
L. p. Tang et al., 2008b: 28, f. 14a-j (f, Dm).

mf qiuae Song & Wang, 1991....................China [urn:lsid:amnh.org:spidersp:030506]
L. qiui Song & Wang, 1991: 1, f. 1A-E (Dmf).
L. qiui Song & Zhu, 1997: 128, f. 91A-D (mf).
L. q. Song, Zhu & Chen, 1999: 482, f. 278E, O (mf).
L. q. Yin et al., 2012: 1286, f. 691a-c (f).

mf saltus Ono, 1979....................Nepal, Bhutan, China [urn:lsid:amnh.org:spidersp:030507]
L. s. Ono, 1979: 99, f. 2-3, 17-20 (Df).
Xysticus himalayaensis Hu & Li, 1987a: 376, f. 40.1-2, 41.1-2 (mf, misidentified).
Xysticus mandali Hu & Li, 1987b: 324, f. 43.4-6 (f, misidentified).
L. s. Song & Zhu, 1997: 129, f. 92A-D (mf).
L. s. Song, Zhu & Chen, 1999: 482, f. 278F, P (mf).
L. s. Hu, 2001: 341, f. 205.1-5 (mf).
L. s. Ono, 2001b: 220, f. 36-43 (mf).
L. s. Tang et al., 2008b: 31, f. 15a-l (mf).

mf silvanus Ono, 1980....................China, Taiwan [urn:lsid:amnh.org:spidersp:030508]
L. s. Ono, 1980a: 212, f. 25-30 (Dmf).
L. s. Song & Chai, 1990: 369, f. 7A-C (f).
L. s. Song & Li, 1997: 425, f. 30A-C (f).
L. s. Song & Zhu, 1997: 130, f. 93A-F (mf).
L. s. Song, Zhu & Chen, 1999: 482, f. 278G, 279E (mf).
L. s. Yin et al., 2012: 1287, f. 692a-c (f).

m sorsogonensis Barrion & Litsinger, 1995....................Philippines [urn:lsid:amnh.org:spidersp:030509]
L. s. Barrion & Litsinger, 1995: 259, f. 151a-d (Dm).

f spirellus Tang et al., 2008....................China [urn:lsid:amnh.org:spidersp:041574]
L. s. Tang et al., 2008b: 31, f. 16a-f (Df).

mf subdianicus Tang et al., 2008....................China [urn:lsid:amnh.org:spidersp:041575]
L. s. Tang et al., 2008b: 32, f. 17a-l (Dmf).

mf suwertikos Barrion & Litsinger, 1995....................Philippines [urn:lsid:amnh.org:spidersp:030510]
L. s. Barrion & Litsinger, 1995: 260, f. 152a-i (Dmf).

m torsivus Zhang, Zhu & Tso, 2006....................China, Taiwan [urn:lsid:amnh.org:spidersp:040722]
L. t. Zhang, Zhu & Tso, 2006b: 79, f. 4-6 (Dm).
L. t. Tang et al., 2008b: 34, f. 18a-e (m).

mf transversus Tang et al., 2008....................China [urn:lsid:amnh.org:spidersp:041576]
L. t. Tang et al., 2008b: 35, f. 19a-l (Dmf).

mf umalii Barrion & Litsinger, 1995....................Philippines [urn:lsid:amnh.org:spidersp:030511]
L. u. Barrion & Litsinger, 1995: 257, f. 150a-i (Dmf).

mf uniprocessus Tang et al., 2008....................China [urn:lsid:amnh.org:spidersp:041577]
L. u. Tang et al., 2008b: 37, f. 20a-j (Dmf).

f wenensis Song, 1995....................China [urn:lsid:amnh.org:spidersp:030512]
L. minimus Tang, Song & Zhu, 1995: 19, f. 4A-B (f, misidentified).
L. w. Song, 1995: 125 (Df).
L. w. Song & Zhu, 1997: 130, f. 94A-B (f).
L. w. Song, Zhu & Chen, 1999: 482, f. 278H (f).

mf wittmeri Ono, 2001....................Bhutan [urn:lsid:amnh.org:spidersp:038186]
L. w. Ono, 2001b: 226, f. 56-61 (Dmf).