The World Spider Catalog, V15 by N. I. Platnick (c) 2000 - 2014 AMNH

Family: THERIDIIDAE

Gen. Phycosoma O. P.-Cambridge, 1879 [urn:lsid:amnh.org:spidergen:03526]

N.B.: removed from the synonymy of Euryopis Menge, 1868 by Fitzgerald & Sirvid, 2003: 28 (contra Levi & Levi, 1962: 26), considered a junior synonym of Trigonobothrys Simon, 1889 by Fitzgerald & Sirvid, 2003: 28 but actually a senior synonym of that name; Trigonobothrys was removed from the synonymy of Dipoena Thorell, 1869 by Yoshida, 2002a: 14, contra Levi & Levi, 1962: 25-32 and was considered by Yoshida, 2002a: 14 to include "the species of the D. lineatipes group designated by Levi (1953)" but only taxa explicitly transferred by him are listed here, until a wider, worldwide sample of taxa can be studied; Trigonobothrys considered part of Lasaeola Simon, 1881 by Wunderlich, 2008b: 283.

In synonymy:
P. coreanum (Paik, 1995, T from Dipoena) = P. martinae (Roberts, 1983) (Yoshida & Ono, 2000: 132, sub Dipoena).
P. decamaculatum (Chen, Peng & Zhao, 1992, T from Dipoena) = P. martinae (Roberts, 1983) (Zhu, 1998: 236, sub Dipoena).
P. dissimilis (O. P.-Cambridge, 1905, T from Dipoena) = P. inornatum (O. P.-Cambridge, 1861) (Bristowe, 1939: 63, sub Dipoena).
P. furtivum (Chickering, 1951, T from Dipoena) = P. altum (Keyserling, 1886) (Levi, 1963a: 159, sub Dipoena).
P. hubeiensis (Chen & Zhao, 1996, T from Dipoena) = P. sinica (Zhu, 1992) (Zhu, 1998: 238, sub Dipoena).
P. immaculatum (Zhu, 1998, T from Dipoena) = P. flavomarginatum (Bösenberg & Strand, 1906) (Yoshida, 2002a: 14, sub Trigonobothrys).
P. luteum (Keyserling, 1891, T from Euryopis) = P. altum Keyserling, 1886 (Levi, 1963a: 159, sub Dipoena).
P. kayaensis (Paik, 1996, T from Dipoena) = P. japonicum (Yoshida, 1985) (Yoshida & Ono, 2000: 128, sub Dipoena japonica, but see Marusik & Koponen, 2000: 65).
P. ruedai (Barrion & Litsinger, 1995, T from Dipoena) = P. martinae (Robert, 1983) (Yoshida & Ono, 2000: 132, sub Dipoena).

mf altum (Keyserling, 1886)....................Mexico to Brazil, Hawaii [urn:lsid:amnh.org:spidersp:007340]
Dipoena alta Keyserling, 1886b: 45, pl. 12, f. 159 (Dm).
Euryopis lutea Keyserling, 1891: 227, pl. 9, f. 168 (Df).
Dipoena pallida Chickering, 1943b: 364, f. 50-51 (Df, preoccupied by Emerton, 1913).
Dipoena furtiva Chickering, in Roewer, 1951: 455 (replacement name).
Dipoena furtiva Levi, 1953a: 3, f. 18 (f).
Dipoena alta Levi, 1963a: 159, f. 138-139 (Sf).
Dipoena alta Roberts, 1979: 201, f. 1-57, 107, 111-114 (mf).
Dipoena alta Coddington, 1990: 17, f. 76 (m).
P. alta Fitzgerald & Sirvid, 2004: 10 (Tmf from Dipoena, after Yoshida, 2002a).
P. a. Ibarra-Núñez, Maya-Morales & Chamé-Vázquez, 2011: 1186, f. 5f (f).

mf amamiense (Yoshida, 1985)....................Russia, China, Korea, Japan, Ryukyu Is. [urn:lsid:amnh.org:spidersp:007341]
Pholcomma amamiensis Yoshida, 1985c: 11, f. 4-5, 11-12 (Df).
Dipoena amamiensis Yoshida, 1991e: 35 (Tf from Pholcomma, Dm now placed in Lasaeola okinawana).
Dipoena japonica Yoshida, 1991e: 33, f. 1, 3-4 (f, misidentified).
Dipoena amamiensis Zhu, 1992: 108, f. 1-4 (f, may be that of D. sticta).
Dipoena amamiensis Paik, 1995c: 33, f. 7-12 (f).
Dipoena amamiensis Zhu, 1998: 242, f. 159A-D (f, may be that of D. sticta, not m).
Dipoena amamiensis Song, Zhu & Chen, 1999: 110, f. 53G-H (f, not m).
Dipoena amamiensis Yoshida & Ono, 2000: 130, f. 7-10 (f, Dm).
Trigonobothrys amamiensis Yoshida, 2002a: 14, f. 6 (Tmf from Dipoena).
Trigonobothrys amamiensis Yoshida, 2003a: 177, f. 489-492 (mf).
P. a. Yoshida, 2009b: 391, f. 335-337 (mf).
Dipoena amamiensis Zhu & Zhang, 2011: 91, f. 51A-D (f).
P. amamiensis Yin et al., 2012: 369, f. 154a-e (mf).

mf corrugum Gao & Li, 2014....................China [urn:lsid:amnh.org:spidersp:046691]
P. c. Gao & Li, 2014: 63, f. 51A-D, 52A-B, 53A-E, 54A-D (Dmf).

m crenatum Gao & Li, 2014....................China [urn:lsid:amnh.org:spidersp:046692]
P. c. Gao & Li, 2014: 70, f. 55A-B, 56A-D, 57A-D (Dm).

m diaoluo Zhang & Zhang, 2012....................China [urn:lsid:amnh.org:spidersp:045000]
P. d. Zhang & Zhang, 2012b: 33, f. 6-8 (Dm).

mf digitula Zhang & Zhang, 2012....................China [urn:lsid:amnh.org:spidersp:044999]
P. d. Zhang & Zhang, 2012b: 31, f. 1-5 (Dmf).

mf excisum (Simon, 1889)....................Madagascar [urn:lsid:amnh.org:spidersp:007381]
Trigonobothrys excisus Simon, 1889b: 231 (Df).
Trigonobothrys excisus Simon, 1894a: 559, f. 571 (f).
Trigonobothrys excisus Levi & Levi, 1962: 41, f. 52-54 (f, Dm).
P. e. Fitzgerald & Sirvid, 2004: 8, f. 1c (m).

mf flavomarginatum (Bösenberg & Strand, 1906)....................China, Korea, Japan [urn:lsid:amnh.org:spidersp:007383]
Dipoena flavomarginata Bösenberg & Strand, 1906: 151, pl. 12, f. 279 (Dmf).
Dipoena flavomarginata Ohno & Yaginuma, 1968: 27, f. 1-5 (f).
Dipoena flavomarginata Arita, 1970: 26, f. 13-14 (f).
Dipoena flavomarginata Yoshida, 1978a: 22, f. 3 (m).
Dipoena flavomarginata Yaginuma, 1986a: 42, f. 23.2 (mf).
Dipoena flavomarginata Chikuni, 1989b: 36, f. 33 (mf).
Dipoena flavomarginata Zhu, 1998: 231, f. 150A-D (mf).
Dipoena immaculata Zhu, 1998: 251, f. 167A-D (Dm).
Dipoena flavomarginata Song, Zhu & Chen, 1999: 110, f. 53K-L, P (mf).
Dipoena immaculata Song, Zhu & Chen, 1999: 112, f. 54M-N (m).
Dipoena flavomarginata Yoshida & Ono, 2000: 136, f. 18-20 (mf).
Trigonobothrys flavomarginatus Yoshida, 2002a: 14, f. 25-28 (Tmf from Dipoena, S).
Dipoena flavomarginata Namkung, 2002: 110, f. 13.28a-b (mf).
Trigonobothrys flavomarginatus Yoshida, 2003a: 181, f. 503-506, 595 (mf).
Trigonobothrys flavomarginatus Namkung, 2003: 112, f. 13.28a-b (mf).
Dipoena flavomarginata Lee et al., 2004: 100, unnumbered f. (f).
P. p. Yoshida, 2009b: 391, f. 343-344 (mf).
Dipoena flavomarginata Zhu & Zhang, 2011: 92, f. 52A-E (mf).
Phycosoma flavomarginata Yin et al., 2012: 370, f. 155a-c (f).

mf hainanensis (Zhu, 1998)....................China, Laos [urn:lsid:amnh.org:spidersp:007393]
Dipoena h. Zhu, 1998: 234, f. 153A-F (Dmf).
Dipoena h. Song, Zhu & Chen, 1999: 110, f. 54C-D. K-L (mf).
P. h. Zhang & Zhang, 2012b: 41 (Tmf from Dipoena).

mf hana (Zhu, 1998)....................China [urn:lsid:amnh.org:spidersp:007394]
Dipoena h. Zhu, 1998: 237, f. 155A-C (Df).
Dipoena h. Song, Zhu & Chen, 1999: 110, f. 54E-F (f).
P. h. Zhang & Zhang, 2012b: 36, f. 15-19 (Tf from Dipoena, Dm).

mf inornatum (O. P.-Cambridge, 1861)....................Europe to Azerbaijan [urn:lsid:amnh.org:spidersp:007399]
Theridion i. O. P.-Cambridge, 1861a: 433 (Dmf).
Theridion i. Blackwall, 1864a: 196, pl. 14, f. 127 (mf).
Euryopis inornata Pavesi, 1873a: 90.
Steatoda inornata L. Koch, 1877b: 140.
Euryopis inornata Bertkau, 1880b: 310, pl. 6, f. 11 (m).
Lasaeola inornata Simon, 1881a: 146.
Lasaeola inornata Chyzer & Kulczyn'ski, 1894: 25, pl. 1, f. 18 (mf).
Lasaeola inornata Bösenberg, 1902: 125, pl. 11, f. 160 (mf).
Lasaeola bösenbergi Strand, 1902: 5 (name for specimens erroneously thought misidentified as D. inornata by Bösenberg, 1902).
Lasaeola dissimilis O. P.-Cambridge, 1905: 49. 58, pl. A, f. 6-8 (Dm).
Lasaeola dissimilis O. P.-Cambridge, 1906a: 75, 84, pl. A, f. 1 (Df).
Dipoena inornata Lessert, 1910b: 117.
Dipoena inornata Simon, 1914a: 278, 301, f. 528 (mf).
Dipoena inornata Wiehle, 1937: 187, f. 186-189 (mf).
Dipoena inornata Bristowe, 1939: 63 (S).
Dipoena inornata Locket & Millidge, 1953: 47, f. 31D-E, 32C (mf).
Dipoena inornata Lehtinen & Kleemola, 1962: 110, f. 15 (f).
Dipoena inornata Miller, 1967: 291, pl. XIV, f. 5-6 (f).
Dipoena inornata Tyschchenko, 1971: 144, f. 373, 376 (mf).
Dipoena inornata Miller, 1971: 186, pl. XXXI, f. 22 (f).
Dipoena inornata Palmgren, 1974b: 11, f. 3.6-7 (mf).
Dipoena inornata Roberts, 1985: 176, f. 78c (mf).
Dipoena inornata Heimer & Nentwig, 1991: 284, f. 767 (mf).
Dipoena inornata Roberts, 1995: 268, f. (mf).
Dipoena inornata Roberts, 1998: 283, f. (mf).
P. inornata Fitzgerald & Sirvid, 2004: 10 (Tmf from Dipoena, after Yoshida, 2002a).
Dipoena inornata Almquist, 2005: 60, f. 86a-e (mf).
Dipoena inornata Le Peru, 2011: 440, f. 680 (mf).
P. i. Kovblyuk, Marusik & Omelko, 2012: 252, f. 35-38 (m).

mf jamesi (Roberts, 1979)....................Jamaica, Panama [urn:lsid:amnh.org:spidersp:007403]
Dipoena j. Roberts, 1979: 204, f. 88-106, 115-116 (Dmf).
P. j. Fitzgerald & Sirvid, 2004: 10 (Tmf from Dipoena).

mf japonicum (Yoshida, 1985)....................Korea, Japan [urn:lsid:amnh.org:spidersp:007404]
Pholcomma j. Yoshida, 1985c: 10, f. 3, 8-10 (Df).
Pholcomma j. Yoshida, 1989b: 318, f. 4M (f).
Dipoena japonica Yoshida, 1991e: 33, f. 1, 4 (Tf from Pholcomma, Dm).
Dipoena kayaensis Paik, 1996e: 42, f. 1-14 (Dmf).
Dipoena japonica Kim & Kim, 1997: 20, f. 1-5 (f).
Dipoena japonica Yoshida & Ono, 2000: 128, f. 1-6 (mf, S, but see Marusik & Koponen, 2000: 65, sub D. amamiensis).
Trigonobothrys japonicus Yoshida, 2002a: 14 (Tmf from Dipoena).
Trigonobothrys japonicus Yoshida, 2003a: 175, f. 483-488 (mf).
P. j. Yoshida, 2009b: 391, f. 332-334 (mf).

mf labialis (Zhu, 1998)....................China [urn:lsid:amnh.org:spidersp:007410]
Dipoena l. Zhu, 1998: 241, f. 158A-D (Df).
Dipoena l. Song, Zhu & Chen, 1999: 112, f. 55A-B (f).
P. l. Zhang & Zhang, 2012b: 38, f. 20-24 (Tf from Dipoena, Dm).

m ligulaceum Gao & Li, 2014....................China [urn:lsid:amnh.org:spidersp:046693]
P. l. Gao & Li, 2014: 70, f. 58A-B, 59A-D, 60A-D (Dm).

mf lineatipes (Bryant, 1933)....................USA to Brazil [urn:lsid:amnh.org:spidersp:007417]
Dipoena l. Bryant, 1933a: 174, pl. 1, f. 7 (Df).
Dipoena l. Levi, 1953b: 12, f. 11-15, 120-121 (f, Dm).
Dipoena alta Levi, 1963a: 159 (S, rejected).
Dipoena l. Roberts, 1979: 202, f. 58-87, 108-110 (mf, removed from S).
P. l. Fitzgerald & Sirvid, 2004: 10 (Tmf from Dipoena, after Yoshida, 2002a).

mf martinae (Roberts, 1983)....................Seychelles, India, China, Korea, Ryukyu Is., Philippines [urn:lsid:amnh.org:spidersp:007423]
Dipoena m. Roberts, 1983: 227, f. 32-35 (Dm).
Dipoena japonica Yoshida, 1991e: 33, f. 2 (f, misidentified).
Dipoena decamaculata Chen, Peng & Zhao, 1992: 270, f. 1-5 (Dmf).
Dipoena coreana Paik, 1995c: 32, f. 1-6 (Dm).
Dipoena ruedai Barrion & Litsinger, 1995: 454, f. 274a-j (Dm).
Dipoena m. Zhu, 1998: 236, f. 154A-F (m, Sf).
Dipoena m. Song, Zhu & Chen, 1999: 112, f. 55C-D, K-L (mf).
Dipoena m. Yoshida & Ono, 2000: 132, f. 11-16 (mf, S).
Trigonobothrys m. Yoshida, 2002a: 14 (Tmf from Dipoena).
Trigonobothrys m. Yoshida, 2003a: 178, f. 493-498 (mf).
Trigonobothrys m. Sudhikumar, Mathew & Sebastian, 2004: 52, f. a-h (mf).
P. m. Saaristo, 2006: 52, f. 1-4 (mf).
P. m. Yoshida, 2009b: 391, f. 338-340 (mf).
P. m. Saaristo, 2010: 240, f. 37.9-12 (mf).

mf menustya (Roberts, 1983)....................Seychelles [urn:lsid:amnh.org:spidersp:007427]
Dipoena m. Roberts, 1983: 224, f. 24-31 (Dmf).
P. m. Fitzgerald & Sirvid, 2004: 10 (Tmf from Dipoena).
P. m. Saaristo, 2010: 239, f. 37.1-8 (mf).

mf mustelinum (Simon, 1889)....................Russia, China, Korea, Japan, Krakatau [urn:lsid:amnh.org:spidersp:007432]
Euryopis mustelina Simon, 1889d: 251 (Df).
Euryopis mustelina Bösenberg & Strand, 1906: 137, pl. 3, f. 2, pl. 12, f. 276 (f).
Euryopis mustelina Paik, 1962: 74, f. 1 (f).
Dipoena mustelina Yaginuma, 1967b: 88, f. 1a-c (Tf from Euryopis, Dm).
Dipoena mustelina Yaginuma, 1986a: 42, f. 23.3 (mf).
Dipoena mustelina Chikuni, 1989b: 36, f. 32 (mf).
Dipoena mustelina Zhu, 1992: 109, f. 5-6 (f).
Dipoena mustelina Zhu, 1998: 240, f. 157A-D (mf).
Dipoena mustelina Song, Zhu & Chen, 1999: 112, f. 55E-F, M (mf).
Dipoena mustelina Yoshida & Ono, 2000: 135, f. 17 (m).
Trigonobothrys mustelinus Yoshida, 2002a: 14 (Tmf from Dipoena).
Dipoena mustelina Namkung, 2002: 111, f. 13.29a-b (mf).
Dipoena mustelina Kim & Cho, 2002: 302, f. 793-798 (mf).
Trigonobothrys mustelinus Yoshida, 2003a: 179, f. 499-502, 594 (mf).
Trigonobothrys mustelinus Namkung, 2003: 113, f. 13.29a-b (mf).
Dipoena mustelina Lee et al., 2004: 100, unnumbered f. (f).
P. m. Yoshida, 2009b: 391, f. 12, 341-342 (mf).
P. mustelina Yin et al., 2012: 372, f. 156a-c (f).

mf nigromaculatum (Yoshida, 1987)....................China, Taiwan, Japan, Ryukyu Is. [urn:lsid:amnh.org:spidersp:007436]
Pholcomma n. Yoshida, 1987c: 29, f. 1-5 (Dmf).
Dipoena nigromaculata Zhu, 1998: 227, f. 147A-E (Tmf from Pholcomma).
Dipoena nigromaculata Song, Zhu & Chen, 1999: 112, f. 55G-H, N (mf).
Trigonobothrys nigromaculatus Yoshida, 2002a: 15 (Tmf from Dipoena).
Trigonobothrys nigromaculatus Yoshida, 2003a: 152, f. 507-511 (mf).
P. n. Yoshida, 2009b: 391, f. 345-347 (mf).

mf oecobioides O. P.-Cambridge, 1879 *....................New Zealand, Chatham Is. [urn:lsid:amnh.org:spidersp:007728]
P. o. O. P.-Cambridge, 1879d: 692, pl. 52, f. 6 (Df).
Trigonobothrys o. Fitzgerald & Sirvid, 2003: 26, f. 1, 2ac (Tf from Euryopis, Dm).
P. o. Fitzgerald & Sirvid, 2004: 10, f. 1a, 2a (mf).

mf sinica (Zhu, 1992)....................China [urn:lsid:amnh.org:spidersp:007486]
Dipoena s. Zhu, 1992: 109, f. 7-13 (Dmf).
Dipoena hubeiensis Chen & Zhao, 1996: 35, f. 1-7 (Dmf).
Dipoena s. Zhu, 1998: 238, f. 156A-G (mf, S).
Dipoena s. Song, Zhu & Chen, 1999: 112, f. 56E-F, K-L (mf).
Dipoena s. Zhu & Zhang, 2011: 93, f. 53A-G (mf).
P. s. Zhang & Zhang, 2012b: 41 (Tmf from Dipoena).
Dipoena s. Yin et al., 2012: 323, f. 126a-d, 13-4g (mf).

mf spundana (Roberts, 1978)....................Seychelles [urn:lsid:amnh.org:spidersp:007487]
Dipoena s. Roberts, 1978: 903, f. 1-4 (Dm).
P. s. Saaristo, 2006: 54, f. 5-12 (Tm from Dipoena, Df).
P. s. Saaristo, 2010: 241, f. 37.13-20 (mf).

mf stellaris (Zhu, 1998)....................China [urn:lsid:amnh.org:spidersp:007489]
Dipoena s. Zhu, 1998: 230, f. 149A-D (Df).
Dipoena s. Song, Zhu & Chen, 1999: 112, f. 56G-H (f).
P. s. Zhang & Zhang, 2012b: 34, f. 9-14 (Tf from Dipoena, Dm).

m stictum (Zhu, 1992)....................China [urn:lsid:amnh.org:spidersp:007490]
Dipoena sticta Zhu, 1992: 110, f. 14-17 (Dm).
Dipoena sticta Zhu, 1998: 252, f. 168A-D (m).
Dipoena sticta Song, Zhu & Chen, 1999: 112, f. 56M-N (m).
P. sticta Yin et al., 2012: 373, f. 157a-c (Tm from Dipoena).

m stigmosum Yin, 2012....................China [urn:lsid:amnh.org:spidersp:046189]
P. stigmosa Yin, in Yin et al., 2012: 374, f. 158a-d (Dm).