The World Spider Catalog, V15 by N. I. Platnick (c) 2000 - 2014 AMNH

Family: AGELENIDAE

Gen. Pireneitega Kishida, 1955 [urn:lsid:amnh.org:spidergen:03778]

P. Kishida, 1955: 11, type Coelotes roscidus L. Koch, 1868=C. segestriformis (Dufour, 1820); N.B.: considered a nomen nudum by Yaginuma, in Brignoli, 1983c: 468 but treated as valid (and a senior synonym of Paracoelotes Brignoli, 1982b: 347, type Coelotes armeniacus Brignoli, 1978) by Wang & Jäger, 2007: 46; see note under Agelenidae regarding familial placement.

Transferred to other genera:
P. wuermlii (Brignoli, 1978) -- see Draconarius.

In synonymy:
P. birulai (Ermolaev, 1927, T from Coelotes) = P. luctuosa (L. Koch, 1878) (Brignoli, 1982b: 350, sub Paracoelotes).
P. csikii (Kulczyn'ski, 1901, T from Coelotes) = P. spinivulva (Simon, 1880) (Song & Hubert, 1983: 5, contra Lehtinen, 1967: 224, sub Paracoelotes).
P. fasciata (Wang et al., 1990, T from Coelotes) = P. spinivulva (Simon, 1881) (Wang, 2002: 116, sub Paracoelotes).
P. japonica (Karsch, 1879, removed from S of Coelotes insidiosus) = P. luctuosa (L. Koch, 1878) (Lehtinen, 1967: 224, sub Coelotes).
P. kekenboschi (de Blauwe, 1973, T from Coelotes) = P. garibaldii (Kritscher, 1969) (Brignoli, 1977f: 17, sub Coelotes).
P. luctuosa schensiensis (Schenkel, 1963, T from Coelotes) = P. spinivulva (Simon, 1880) (Schenkel, 1963: 293, footnote, sub Coelotes csikii).
P. moellendorffi (Karsch, 1881, T from Coelotes) = P. spinivulva (Simon, 1881) (Wang, 2002: 116, sub Paracoelotes).
P. qinlingensis (Peng, Gong & Kim, 1996, T from Coelotes) = P. spinivulva (Simon, 1881) (Wang, 2002: 116, sub Paracoelotes).
P. sakhalinensis (Marusik & Logunov, 1991, T from Coelotes) = P. luctuosa (L. Koch, 1878) (Ovtchinnikov, 1999: 79, sub Paracoelotes birulai).
P. subluctuosa (Zhu & Wang, 1994) = P. taishanensis (Wang et al., 1990) (Wang, 2002: 117, contra Marusik & Koponen, 2000: 56, who placed that name as a junior synonym of P. spinivulva instead, sub Paracoelotes).
P. vulgaris (Paik, 1971, T from Coras) = P. spinivulva (Simon, 1880) (Brignoli, 1982b: 349, sub Paracoelotes csikii).

mf armeniaca (Brignoli, 1978)....................Turkey [urn:lsid:amnh.org:spidersp:023018]
Coelotes armeniacus Brignoli, 1978g: 527, f. 117-123 (Dmf).
Paracoelotes armeniacus Brignoli, 1982b: 347 (Tmf from Coelotes).
Paracoelotes armeniacus Wang, 2002: 118, f. 323-335 (mf).

f bidens (Caporiacco, 1935)....................Karakorum [urn:lsid:amnh.org:spidersp:023019]
Coelotes b. Caporiacco, 1935b: 227, pl. 6, f. 10 (Df).
Coelotes spinivulva Lehtinen, 1967: 224 (S, rejected).
Coelotes b. Brignoli, 1976d: 234, f. 7, 10 (f, removed from S, presumably a close relative of P. spinivulvus).

f cottarellii (Brignoli, 1978)....................Turkey [urn:lsid:amnh.org:spidersp:022746]
Coelotes c. Brignoli, 1978g: 532, f. 130-131 (Df).
Paracoelotes c. Wang, 2002: 114 (Tf from Coelotes).

f fedotovi (Charitonov, 1946)....................Uzbekistan [urn:lsid:amnh.org:spidersp:023020]
Coelotes f. Charitonov, 1946: 20, f. 4 (Df; see also Charitonov, 1969: 84).
Paracoelotes f. Brignoli, 1982b: 350 (Tf from Coelotes).

mf garibaldii (Kritscher, 1969)....................Italy [urn:lsid:amnh.org:spidersp:023021]
Coelotes g. Kritscher, 1969: 292, f. 5-6 (Df).
Coelotes segestriformis de Blauwe, 1973a: 95, f. 90-91 (f, S, rejected).
Coelotes kekenboschi de Blauwe, 1973a: 50, f. 44-45 (Df).
Coelotes g. Brignoli, 1977f: 17, f. 1-3 (f, Dm, S).
Paracoelotes g. Brignoli, 1982b: 349 (Tmf from Coelotes).
Paracoelotes g. Trotta, 2005: 160, f. 199-200 (mf).
P. g. IJland, van Helsdingen & Miller, 2012: 5, f. 3 (m).
P. g. Kovblyuk et al., 2013a: 68, f. 12-16, 28, 32, 36, 40, 43 (mf).

mf involuta (Wang et al., 1990)....................China [urn:lsid:amnh.org:spidersp:022786]
Coelotes involutus Wang et al., 1990: 182, f. 13-19 (Dmf).
Coelotes involutus Song, Zhu & Chen, 1999: 376, f. 220H-I, 222I, 223L (mf).
Paracoelotes involutus Wang, 2002: 114 (Tmf from Coelotes).

f liansui (Bao & Yin, 2004)....................China [urn:lsid:amnh.org:spidersp:040278]
Coelotes l. Bao & Yin, 2004: 455, f. 1-3 (Df).
P. i. Wang & Jäger, 2007: 46 (Tf from Coelotes).
Paracoelotes l. Yin et al., 2012: 1020, f. 528a-c (f).

mf luctuosa (L. Koch, 1878)....................Russia, Central Asia, China, Korea, Japan [urn:lsid:amnh.org:spidersp:023022]
Coelotes luctuosus L. Koch, 1878c: 752, pl. 15, f. 14-16 (Dmf).
Coelotes japonicus Karsch, 1879g: 97, pl. 1, f. 16 (Df).
Coras luctuosus Bösenberg & Strand, 1906: 300, pl. 16, f. 459 (mf).
Coras luctuosus Strand, 1918: 84, pl. 1, f. 18 (f).
Coelotes birulai Ermolajev, 1927: 347, f. 1, 3-5 (Dmf).
Coras luctuosus Saito, 1939: 62, f. 8.2 (f).
Coras corasides Komatsu, 1942a: 1, pl. 1 (Tf from Tegenaria, misidentified per Yaginuma, 1972b: 46).
Coras luctuosus Yaginuma, 1957c: 17, f. 1 (f).
Coras luctuosus Saito, 1959: 43, f. 16a-c (f).
Coras luctuosus Yaginuma, 1960: 93, f. 82.5 (f).
Coelotes luctuosus Lehtinen, 1967: 224 (S).
Coras luctuosus Shinkai, 1969: 34, f. 19 (m).
Coras luctuosus Yaginuma, 1971: 93, f. 82.5 (f).
Coras luctuosus Paik, 1971a: 12, f. 13-21 (mf).
Coelotes luctuosus Nishikawa, 1974: 176, f. 15-16 (mf).
Coelotes luctuosus Shinkai, 1978: 95, f. 38 (m).
Coras luctuosus Paik, 1978e: 354, f. 159.1-3 (mf).
Paracoelotes luctuosus Brignoli, 1982b: 350, f. 9-10 (Tmf from Coras, S).
Coelotes luctuosus Yin, Wang & Hu, 1983: 34, f. 5E (m).
Coras luctuosus Hu, 1984: 206, f. 217.1-3 (mf).
Coras luctuosus Guo, 1985: 111, f. 2-52.1-2 (f).
Coelotes luctuosus Zhu et al., 1985: 123, f. 108a-c (f).
Coelotes luctuosus Yaginuma, 1986a: 147, f. 82.1 (mf).
Coelotes luctuosus Song, 1987: 191, f. 150 (mf).
Coelotes luctuosus Zhang, 1987: 136, f. 113.1-4 (mf).
Coelotes luctuosus Chikuni, 1989b: 100, f. 8 (mf).
Coelotes luctuosus Feng, 1990: 141, f. 116.1-3 (mf).
Coelotes luctuosus Chen & Gao, 1990: 116, f. 146a-b (mf).
Coelotes luctuosus Chen & Zhang, 1991: 187, f. 184.1-3 (mf).
Coelotes sakhalinensis Marusik & Logunov, 1991: 87, f. 1.1-2 (Dm).
Coelotes luctuosus Zhao, 1993: 303, f. 138a-c (mf).
Paracoelotes birulai Ovtchinnikov, 1999: 79 (S).
Paracoelotes luctuosus Song, Zhu & Chen, 1999: 389, f. 229O-P, U (mf).
Paracoelotes luctuosus Song, Zhu & Chen, 2001: 295, f. 188A-C (mf).
Coelotes luctuosus Yoo & Kim, 2002: 26, f. 30 (m).
Paracoelotes luctuosus Namkung, 2002: 402, f. 28.16a-b (mf).
Paracoelotes luctuosus Namkung, 2003: 404, f. 28.16a-b (mf).
P. l. Okumura et al., 2009: 178, f. 44-45 (mf).
P. l. Zhu & Zhang, 2011: 326, f. 236A-E (mf).
Paracoelotes luctuosus Yin et al., 2012: 1021, f. 529a-e, 3-13p-q (mf).

mf luniformis (Zhu & Wang, 1994)....................China [urn:lsid:amnh.org:spidersp:022803]
Coelotes l. Zhu & Wang, 1994: 38, f. 5-8 (Dmf).
Coelotes l. Song, Zhu & Chen, 1999: 376, f. 220P-Q, 222J, 223M (mf).
Paracoelotes l. Wang, 2002: 114 (Tmf from Coelotes).

f major (Kroneberg, 1875)....................Uzbekistan, Tajikistan, China [urn:lsid:amnh.org:spidersp:023023]
Coelotes m. Kroneberg, 1875: 15, pl. 1, f. 6 (Df).
Coelotes m. Schenkel, 1936b: 284, f. 97 (f).
Coelotes m. Charitonov, 1946: 20, f. 5 (f).
Paracoelotes m. Ovtchinnikov, 1988: 142 (Tf from Coelotes).
Coelotes m. Hu & Wu, 1989: 180, f. 150.1-2 (f).
Paracoelotes m. Song, Zhu & Chen, 1999: 389, f. 229Q-R (f).

f neglecta (Hu, 2001)....................China [urn:lsid:amnh.org:spidersp:037813]
Coelotes neglectum Hu, 2001: 139, f. 50.1-2 (Df).
P. n. Wang & Jäger, 2007: 46 (Tf from Coelotes).

mf ovtchinnikovi Kovblyuk et al., 2013....................Russia, Abkhazia, Georgia [urn:lsid:amnh.org:spidersp:045832]
P. o. Kovblyuk et al., 2013a: 64, f. 1-6, 22-26, 30, 34, 38, 42, 48 (Dmf).

mf pyrenaea (Simon, 1870)....................Spain, France [urn:lsid:amnh.org:spidersp:023024]
Coelotes pyrenaeus Simon, 1870b: 293 (Dm).
Coelotes pyrenaeus Simon, 1873a: 169, p. 1, f. 22 (m).
Coelotes pyrenaeus Simon, 1875a: 40, pl. 5, f. 12 (m, Df).
Amaurobius obesus Kulczyn'ski, 1906b: 424, pl. 14, f. 1 (f, misidentified).
Amaurobius pyrenaeus Kulczyn'ski, 1906b: 428, pl. 14, f. 4 (f).
Coelotes pyrenaeus Simon, 1937: 985, 988, 1034, f. 1509, 1517 (mf).
Coelotes pyrenaeus de Blauwe, 1973a: 89, f. 79-82 (mf).
Paracoelotes pyrenaeus Brignoli, 1982b: 349 (Tmf from Coelotes).

mf segestriformis (Dufour, 1820) *....................Europe, Russia [urn:lsid:amnh.org:spidersp:023025]
Drassus s. Dufour, 1820d: 297, pl. 95, f. 1 (Dmf).
Amaurobius roscidus C. L. Koch, 1834: 15 (D).
Clubiona roscida Walckenaer, 1841: 480 (Df).
Amaurobius roscidus C. L. Koch, 1843: 113, f. 829 (f).
Drassus s. Dufour, 1858: 260, pl. 6, f. 2 (mf).
Coelotes roscidus L. Koch, 1868: 40, pl. 2, f. 19 (f).
Coelotes atramentarius Simon, 1875a: 46 (Df).
Coelotes leveillei Simon, 1876c: 92 (Dmf).
Amaurobius obesus Kulczyn'ski, 1906b: 425, pl. 14, f. 25, pl. 15, f. 37, 53, 68 (m, misidentified).
Amaurobius leveillei Kulczyn'ski, 1906b: 426, pl. 15, f. 38, 54, 67 (m).
Amaurobius atramentarius Kulczyn'ski, 1906b: 430, pl. 14, f. 22 (f).
Amaurobius dubius Kulczyn'ski, 1906b: 432, pl. 15, f. 19 (Df).
Coelotes s. Simon, 1937: 895, 988, 1034, f. 1510, 1518 (mf).
Coelotes s. de Blauwe, 1973a: 95, f. 85-89 (mf).
Coelotes s. Brignoli, 1978e: 280, f. 12-14 (mf).
Paracoelotes s. Brignoli, 1982b: 349 (Tmf from Coelotes).
P. s. Kovblyuk et al., 2013a: 68, f. 7-11, 27, 31, 35, 39 (mf).

mf spasskyi (Charitonov, 1946)....................Russia, Georgia, Azerbaijan [urn:lsid:amnh.org:spidersp:023026]
Coelotes s. Charitonov, 1946: 20, f. 6 (Df; see also Charitonov, 1969: 83).
Paracoelotes s. Brignoli, 1982b: 349 (Tf from Coelotes).
Coelotes s. Mcheidze, 1997: 204, f. 429-430 (f).
P. s. Kovblyuk et al., 2013a: 71, f. 17-21, 29, 33, 37, 41, 44 (f, Dm).

mf spinivulva (Simon, 1880)....................Russia, China, Korea [urn:lsid:amnh.org:spidersp:023027]
Coelotes s. Simon, 1880b: 116, pl. 3, f. 16 (Df).
Cedicus möllendorffi Karsch, 1881f: 220 (Df).
Coelotes csikii Kulczyn'ski, 1901b: 341, pl. 13, f. 14-15 (Dm).
Coelotes caraftensis Kishida, 1924: 518 (Dj; N.B.: considered a nomen dubium by Marusik et al., 1993b: 82).
Coelotes csikii Ermolajev, 1927: 347, f. 2 (m).
Coelotes laticeps Schenkel, 1936b: 181, f. 60c (f, misidentified).
Coelotes luctuosus schensiensis Schenkel, 1963: 293, f. 165a-c (Dm).
Coelotes csikii Schenkel, 1963: 293, footnote (S).
Coelotes s. Lehtinen, 1967: 224 (S, rejected).
Coelotes möllendorffi Lehtinen, 1967: 448, f. 240 (f).
Coras vulgaris Paik, 1971a: 8, f. 1-12 (Dmf).
Coras vulgaris Paik, 1978e: 356, f. 160.1-3 (mf).
Paracoelotes csikii Brignoli, 1982b: 349 (Tmf from Coelotes, S).
Coelotes s. Song & Hubert, 1983: 5, f. 10-12 (mf, S).
Coelotes s. Zhu et al., 1985: 123, f. 109a-c (m).
Coelotes s. Song, 1987: 194, f. 153 (mf).
Coelotes s. Zhang, 1987: 137, f. 114.1-3 (mf).
Coelotes s. Hu & Wu, 1989: 179, f. 149.1-3 (mf).
Coelotes s. Feng, 1990: 142, f. 117.1-3 (m).
Coelotes fasciatus Wang et al., 1990: 180, f. 6-12 (Dmf).
Coelotes qinlingensis Peng, Gong & Kim, 1996: 19, f. 10-14 (Dmf).
Coelotes fasciatus Song, Zhu & Chen, 1999: 375, f. 217O-P, 222A, 223F (mf).
Coelotes qinlingensis Song, Zhu & Chen, 1999: 377, f. 221A'-B', 223E, 224G (mf).
Paracoelotes s. Song, Zhu & Chen, 1999: 389, f. 229S-T, V (mf).
Paracoelotes s. Marusik & Koponen, 2000: 56 (S, rejected).
Coelotes s. Hu, 2001: 147, f. 56.1-3 (mf).
Paracoelotes s. Song, Zhu & Chen, 2001: 296, f. 189A-C (mf).
Paracoelotes spinivulvus Wang, 2002: 116 (S).
Coelotes vulgaris Yoo & Kim, 2002: 30, f. 75-76 (m).
Paracoelotes s. Namkung, 2002: 401, f. 28.15a-b (mf).
Paracoelotes s. Kim & Cho, 2002: 186, f. 373-378 (mf).
Paracoelotes s. Namkung, 2003: 403, f. 28.15a-b (mf).
P. s. Wang & Jäger, 2010: 1170, f. 1E (f).
P. s. Zhu & Zhang, 2011: 327, f. 237A-F (mf).
Paracoelotes s. Yin et al., 2012: 1023, f. 530a-c (mf).

mf taishanensis (Wang et al., 1990)....................China [urn:lsid:amnh.org:spidersp:022883]
Coelotes t. Wang et al., 1990: 178, f. 1-5 (Dmf).
Coelotes subluctuosus Zhu & Wang, 1994: 39, f. 9-12 (Dmf).
Coelotes subluctuosus Song, Zhu & Chen, 1999: 378, f. 225E-F, 227B, 228E (mf).
Coelotes t. Song, Zhu & Chen, 1999: 378, f. 225I-J, 227D, 228G (mf).
Paracoelotes t. Wang, 2002: 117 (Tmf from Coelotes, S).

mf taiwanensis Wang & Ono, 1998....................Taiwan [urn:lsid:amnh.org:spidersp:023028]
Paracoelotes t. Wang & Ono, 1998: 155, f. 31-36 (Dmf).

mf tianchiensis (Wang et al., 1990)....................China [urn:lsid:amnh.org:spidersp:022891]
Coelotes t. Wang et al., 1990: 212, f. 81-85 (Dmf).
Coelotes t. Song, Zhu & Chen, 1999: 378, f. 225Q-R, 227H, 228I (mf).
Paracoelotes t. Wang, 2002: 118 (Tmf from Coelotes).

mf triglochinata (Zhu & Wang, 1991)....................China [urn:lsid:amnh.org:spidersp:022896]
Coelotes triglochinatus Zhu & Wang, 1991: 1, f. 1-4 (Dmf).
Coelotes triglochinatus Song, Zhu & Chen, 1999: 388, f. 225W-X, 227J, 228K (mf).
P. t. Wang & Jäger, 2007: 48 (Tf from Coelotes, m probably mismatched, possibly a Draconarius species).

mf xinping Zhang, Zhu & Song, 2002....................China [urn:lsid:amnh.org:spidersp:037841]
Paracoelotes x. Zhang, Zhu & Song, 2002: 54, f. 7-10 (Dmf).
P. x. Zhu & Zhang, 2011: 329, f. 238A-E (mf).