The World Spider Catalog, V15 by N. I. Platnick (c) 2000 - 2014 AMNH

Family: GNAPHOSIDAE

Gen. Trachyzelotes Lohmander, 1944 [urn:lsid:amnh.org:spidergen:02490]

N.B.: described as a subgenus of Zelotes Gistel, 1848 by Lohmander, 1944: 13, type Zelotes pedestris (C. L. Koch, 1837); elevated to genus and considered a senior synonym of Simonizelotes Marinaro, 1967: 702, type undesignated, apparently intended to be Zelotes barbatus (L. Koch, 1866) by Platnick & Murphy, 1984: 3.

Transferred to other genera:
T. manytchensis Ponomarev & Tsvetkov, 2006 -- see Zelotes.

In synonymy:
T. acanthognatha (Purcell, 1907, T from Camillina) = T. jaxartensis (Kroneberg, 1875) (Platnick & Murphy, 1984: 10).
T. agilis (Bryant, 1936, T from Drassyllus) = T. lyonneti (Audouin, 1826) (Platnick & Murphy, 1984: 6).
T. barbaranus (Chamberlin, 1922, T from Nodocion) = T. jaxartensis (Kroneberg, 1875) (Ubick & Roth, 1973a: 3, sub Drassyllus peninsulanus).
T. brasilianus (Keyserling, 1891, T from Zelotes) = T. lyonneti (Audouin, 1826) (Platnick & Murphy, 1984: 6).
T. cavaleriei (Schenkel, 1963, T from Zelotes) = T. jaxartensis (Kroneberg, 1875) (Platnick & Murphy, 1984: 11).
T. chamberlini (Roewer, 1951, T from Nodocion) = T. lyonneti (Audouin, 1826) (Ubick & Roth, 1973a: 2, sub Drassyllus agilis).
T. chohanius (Patel & Patel, 1975, T from Scotophaeus) = T. jaxartensis (Kroneberg, 1875) (Platnick & Murphy, 1984: 11).
T. costatus (Denis, 1952) = T. bardiae (Caporiacco, 1928) (Levy, 1998c: 110).
T. errans (Benoit, 1977, T from Zelotes) = T. lyonneti (Audouin, 1826) (Platnick & Murphy, 1984: 7).
T. foveolatus (Simon, 1880, T from Zelotes) = T. jaxartensis (Kroneberg, 1875) (Platnick & Song, 1986: 18).
T. indraprastha (Tikader & Gajbe, 1975, T from Drassodes) = T. jaxartensis (Kroneberg, 1875) (Platnick & Murphy, 1984: 11).
T. insipiens (Simon, 1885, T from Zelotes) = T. jaxartensis (Kroneberg, 1875) (Platnick & Murphy, 1984: 10).
T. iugans (Chamberlin, 1922, T from Nodocion) = T. jaxartensis (Kroneberg, 1875) (Ubick & Roth, 1973a: 3, sub Drassyllus peninsulanus).
T. microbarbatus (Marinaro, 1967, T from Zelotes) = T. mutabilis (Simon, 1878) (Platnick & Murphy, 1984: 18).
T. peninsulanus (Banks, 1898, T from Drassyllus) = T. jaxartensis (Kroneberg, 1875) (Platnick & Murphy, 1984: 10).
T. philippsoni (Denis, 1956, T from Zelotes) = T. lyonneti (Audouin, 1826) (Platnick & Murphy, 1984: 7).
T. postrema (Tucker, 1923, T from Camillina) = T. jaxartensis (Kroneberg, 1875) (Platnick, 1997: 97).
T. rubicundulus (Simon, 1878, T from Zelotes) = T. fuscipes (L. Koch, 1866) (Platnick & Murphy, 1984: 15).
T. samoensis (Berland, 1934, T from Zelotes) = T. kulczynskii (Bösenberg, 1902) (Platnick & Murphy, 1984: 13).
T. solaster (Chamberlin, 1937, T from Nodocion, where omitted by Roewer) = T. lyonneti (Audouin, 1826) (Ubick & Roth, 1973a: 2, sub Drassyllus agilis).
T. sorex (Denis, 1945, T from Zelotes) = T. jaxartensis (Kroneberg, 1875) (Platnick & Murphy, 1984: 11).
T. spinibarbis (Simon, 1897, T from Camillina) = T. jaxartensis (Kroneberg, 1875) (Platnick & Murphy, 1984: 10).
T. ursinus (O. P.-Cambridge, 1872, T from Zelotes) = T. lyonneti (Audouin, 1826) (Platnick & Murphy, 1984: 6).
T. zagistus (Ponomarev, 1981) = T. adriaticus (Caporiacco, 1951) (Ponomarev & Tsvetkov, 2004: 94).

mf adriaticus (Caporiacco, 1951)....................Italy to China [urn:lsid:amnh.org:spidersp:028023]
Zelotes a. Caporiacco, 1951a: 87, f. 8a-b (Dm).
Zelotes zagistus Ponomarev, 1981: 62, f. 4-5 (Dm; palpal illustrations and noted relationship to T. barbatus leave no doubt the species belongs here rather than in Zelotes) [urn:lsid:amnh.org:spidersp:028038].
T. a. Platnick & Murphy, 1984: 9, f. 11-14 (Tm from Zelotes, Df).
T. a. Hu & Wu, 1989: 283, f. 228.1-2 (f).
T. a. Song, Zhu & Chen, 1999: 455, f. 265E, O (mf).
T. a. Tuneva & Esyunin, 2002: 223, f. 30-34 (mf).
T. a. Chatzaki, Thaler & Mylonas, 2003: 54, f. 30-35, 38-39 (f, not m, S).
T. a. Song, Zhu & Zhang, 2004: 232, f. 135A-D (mf).
T. a. Ponomarev & Tsvetkov, 2004: 94 (S).
T. a. Chatzaki, 2010b: 45, f. 2-5, 13 (mf).

f baiyuensis Xu, 1991....................China [urn:lsid:amnh.org:spidersp:040934]
T. b. Xu, 1991b: 39, f. 22-23 (Df).
T. b. Song, Zhu & Zhang, 2004: 233, f. 136A-B (f).

mf barbatus (L. Koch, 1866)....................Mediterranean to Central Asia, USA [urn:lsid:amnh.org:spidersp:028024]
Melanophora barbata L. Koch, 1866: 161, pl. 6, f. 101-103 (Dmf).
Prosthesima barbata Simon, 1878a: 45.
Prosthesima barbata Chyzer & Kulczyn'ski, 1897: 208, pl. 7, f. 46, pl. 8, f. 23 (mf).
Zelotes b. Simon, 1914a: 153, 213, f. 282-283 (mf).
Zelotes b. Denis, 1952f: 115, f. 1-3 (mf).
Zelotes b. Jézéquel, 1962a: 522, f. 3 (f).
T. b. Platnick & Murphy, 1984: 15, f. 27-30 (Tmf from Zelotes).
Zelotes b. Pesarini, 2000: 385, f. 18-19 (mf).
T. b. Chatzaki, Thaler & Mylonas, 2003: 53, f. 20-21, 26-27 (mf).
T. b. Kovblyuk & Tuneva, 2009: 157, f. 10-16 (mf).

mf bardiae (Caporiacco, 1928)....................Mediterranean [urn:lsid:amnh.org:spidersp:028025]
Zelotes b. Caporiacco, 1928b: 87, f. 4 (Dm).
Zelotes costatus Denis, 1952f: 118, f. 4 (Df).
T. costatus Platnick & Murphy, 1984: 20, f. 43-46 (Tf from Zelotes, Dm).
T. b. Levy, 1998c: 110, f. 31-32 (Tm from Zelotes, Sf).

mf chybyndensis Tuneva & Esyunin, 2002....................Russia [urn:lsid:amnh.org:spidersp:038081]
T. c. Tuneva & Esyunin, 2002: 224, f. 35-38 (Dmf).

mf cumensis (Ponomarev, 1979)....................Russia [urn:lsid:amnh.org:spidersp:028170]
Zelotes c. Ponomarev, 1979a: 923, f. 8 (Df).
T. c. Ponomarev & Tsvetkov, 2004: 94, f. 11-12 (Tf from Zelotes, Dm).
T. c. Ponomarev, 2008b: 82, f. 4 (m).

mf fuscipes (L. Koch, 1866)....................Mediterranean, China [urn:lsid:amnh.org:spidersp:028026]
Melanophora f. L. Koch, 1866: 189, pl. 8, f. 127-129 (Dmf).
Prosthesima f. Simon, 1878a: 51.
Prosthesima rubicundula Simon, 1878a: 89 (Dmf).
Zelotes rubicundulus Simon, 1914a: 159, 166, 216, f. 311, 340 (mf).
Zelotes f. Simon, 1914a: 175, 216, f. 365 (f).
Zelotes rubicundulus Jézéquel, 1962a: 527, f. 11 (f).
Zelotes rubicundulus Miller, 1967: 272, pl. III, f. 19 (m).
T. f. Platnick & Murphy, 1984: 15, f. 31-34 (Tmf from Zelotes, S).
T. f. Hu & Wu, 1989: 284, f. 229.1-4 (mf).
T. f. Song, Zhu & Chen, 1999: 455, f. 265F, P (mf).
T. f. Song, Zhu & Zhang, 2004: 234, f. 137A-D (mf).

m glossus (Strand, 1915)....................Turkey, Israel [urn:lsid:amnh.org:spidersp:028027]
Prosthesima glossa Strand, 1915c: 140 (Dm).
T. g. Levy, 1998c: 109, f. 29-30 (Tm from Zelotes).

mf holosericeus (Simon, 1878)....................Mediterranean [urn:lsid:amnh.org:spidersp:028028]
Prosthesima holosericea Simon, 1878a: 47, pl. 14, f. 10 (Dmf).
Zelotes h. Simon, 1914a: 153, 213, f. 280-281 (mf).
Zelotes h. Jézéquel, 1962a: 522, f. 2 (f).
T. h. Platnick & Murphy, 1984: 17, f. 35-38 (Tmf from Zelotes).

mf huberti Platnick & Murphy, 1984....................Algeria, Italy [urn:lsid:amnh.org:spidersp:028029]
T. h. Platnick & Murphy, 1984: 21, f. 47-50 (Dmf).
T. h. IJland, van Helsdingen & Miller, 2012: 8, f. 8a-c (m).

mf jaxartensis (Kroneberg, 1875)....................Holarctic, Senegal, South Africa, Hawaii [urn:lsid:amnh.org:spidersp:028030]
Melanophora j. Kroneberg, 1875: 23, pl. 2, f. 1 (Df).
Prosthesima foveolata Simon, 1880b: 117, pl. 3, f. 17 (Df).
Prosthesima insipiens Simon, 1886g: 382 (Dmf).
Echemus spinibarbis Simon, 1897h: 97 (Df).
Prosthesima peninsulana Banks, 1898b: 217, pl. 13, f. 16 (Df).
Prosthesima iaxartensis Kulczyn'ski, 1901b: 323.
Melanophora acanthognathus Purcell, 1907: 333, pl. 15, f. 66 (Df).
Zelotes peninsulanus Petrunkevitch, 1911: 150.
Nodocion barbaranus Chamberlin, 1922: 154 (Df).
Nodocion iugans Chamberlin, 1922: 154 (Dm).
Camillina acanthognathus Tucker, 1923: 335.
Camillina postrema Tucker, 1923: 344, pl. 9, f. 61 (Dm).
Nodocion barbaranus Chamberlin, 1936a: 12, f. 16 (f).
Zelotes sorex Denis, 1945b: 46, f. 12 (Df).
Zelotes cavaleriei Schenkel, 1963: 50, f. 25a-b (Df).
Drassyllus peninsulanus Ubick & Roth, 1973a: 3 (Tf from Zelotes, Sm).
Scotophaeus chohanius Patel & Patel, 1975a: 35, f. 2a-e (Df).
Drassodes indraprastha Tikader & Gajbe, 1975: 276, f. 11-14 (Df).
Drassodes indraprastha Tikader, 1982b: 419, f. 283-287 (f).
Zelotes cavaleriei Hayashi, 1983a: 11, f. 11-15 (f).
Zelotes cavaleriei Zhang & Zhu, 1983: 166, f. 4-6 (mf).
Zelotes cavalierei Brignoli, 1983c: 578.
Zelotes cavaleriei Hu, 1984: 282, f. 297.1-4 (mf).
T. j. Platnick & Murphy, 1984: 10, f. 19-22 (Tmf from Zelotes, S).
Zelotes cavaleriei Guo, 1985: 145, f. 1-3 (mf).
Zelotes cavaleriei Zhu et al., 1985: 154, f. 137a-c (f).
T. j. Platnick & Song, 1986: 18, f. 71-74 (mf, S).
T. j. Paik, 1986d: 33, f. 44-52 (mf).
Zelotes j. Yaginuma, 1986a: 192, f. 106.7 (f).
T. j. Song, 1987: 343, f. 297 (mf).
Zelotes cavaleriei Zhang, 1987: 186, f. 158.1-4 (mf).
Zelotes j. Chikuni, 1989b: 120, f. 11 (mf).
T. j. Hu & Wu, 1989: 286, f. 228.3-4 (f).
T. j. Chen & Zhang, 1991: 238, f. 248.1-4 (mf).
T. j. Xu, 1991b: 39, f. 24 (f).
Zelotes cavaleriei Zhao, 1993: 127, f. 61a-c (mf).
Zelotes cavaleriei Barrion & Litsinger, 1995: 184, f. 105a-b (f, in part; not m, f. 106a-g, which belongs to Setaphis).
T. j. Platnick, 1997: 97 (S).
T. j. Levy, 1998c: 107, f. 25-28 (mf).
T. j. Song, Zhu & Chen, 1999: 455, f. 265G, Q (mf).
T. j. Song, Zhu & Chen, 2001: 357, f. 233A-D (mf).
T. j. Namkung, 2002: 484, f. 37.25a-b (mf).
T. j. Namkung, 2003: 487, f. 37.25a-b (mf).
T. j. Song, Zhu & Zhang, 2004: 236, f. 138A-D (mf).
T. j. Jung et al., 2005: 196, f. 38, 76, 97, 136-137 (mf).
T. j. Jocqué & Dippenaar-Schoeman, 2006: 128, f. 41i-k (mf).
T. j. Kamura, 2009: 493, f. 150-152 (mf).
T. j. Zhu & Zhang, 2011: 403, f. 287A-D (mf).
T. j. Yin et al., 2012: 1204, f. 642a-e (mf).

mf kulczynskii (Bösenberg, 1902)....................Macedonia, Japan, Caribbean, Colombia, Brazil, Samoa [urn:lsid:amnh.org:spidersp:028031]
Prosthesima k. Bösenberg, 1902: 313, pl. 29, f. 463 (Df).
Zelotes samoensis Berland, 1934b: 325, f. 7-8 (Df).
T. k. Platnick & Murphy, 1984: 13, f. 23-26 (Tf from Zelotes, Dm, S, removed from S of T. barbatus contra Braun, 1981: 377).
T. k. Kamura, 1997: 79, f. 5-8 (mf).
T. k. Brescovit, Freitas & Vasconcelos, 2008: 329, f. 3-6 (mf).
T. k. Kamura, 2009: 493, f. 153-155 (mf).

mf lyonneti (Audouin, 1826)....................Mediterranean to Central Asia, USA to Peru, Brazil [urn:lsid:amnh.org:spidersp:028032]
Drassus lyonetii Audouin, 1826: 383, pl. 5, f. 6 (Dm).
Melanophora ursina O. P.-Cambridge, 1872a: 245, pl. 16, f. 25 (Df).
Drassus secretus Thorell, 1875c: 92 (Df).
Prosthesima l. Simon, 1878a: 98.
Prosthesima ursina Simon, 1878a: 99.
Prosthesima oceanica Simon, 1883: 273, pl. 8, f. 6 (Df).
Drassus brasilianus Keyserling, 1891: 34, pl. 1, f. 13 (Df).
Prosthesima l. Chyzer & Kulczyn'ski, 1897: 208, pl. 7, f. 34, 45 (mf).
Latonigena brasiliana Mello-Leitão, 1918a: 72, f. 42 (Dm).
Nodocion agilis Bryant, 1936a: 93, pl. 3, f. 3 (Dm).
Nodocion zelotoides Chamberlin, 1936b: 6, f. 20 (Df; preoccupied by Worley, 1928).
Nodocion solaster Chamberlin, 1937c: 170 (replacement name for N. zelotoides).
Zelotes brasilianus Mello-Leitão, 1943a: 216, f. 42 (mf).
Nodocion chamberlini Roewer, 1951: 443 (superfluous replacement name for Nodocion zelotoides).
Zelotes philippsoni Denis, 1956b: 275, f. 1 (Df).
Zelotes l. Denis, 1962a: 38, f. 26-28 (mf).
Drassyllus agilis Ubick & Roth, 1973a: 2 (Tm from Nodocion, Sf).
Zelotes errans Benoit, 1977c: 55, f. 19a-f (Df).
T. l. Platnick & Murphy, 1984: 6, f. 7-10 (Tmf from Zelotes, S).
T. l. Levy, 1998c: 105, f. 19-22 (mf).
T. l. Tuneva, 2001: 1521, f. 1-2 (f).
T. l. Chatzaki, Thaler & Mylonas, 2003: 54, f. 24-25, 29 (mf).
T. l. Senglet, 2004: 101, f. 42, 45-46 (mf).
T. l. Kovblyuk, 2005a: 141, f. 5-9 (mf).
T. l. Jiménez, Bizuet-Flores & Corcuera-Martínez del Río, 2012: 145, f. 1-3 (mf).

mf malkini Platnick & Murphy, 1984....................Romania, Crete, Turkey, Ukraine [urn:lsid:amnh.org:spidersp:028033]
Zelotes barbatus Miller, 1967: 271, pl. III, f. 17-18, pl. VI, f. 5 (mf, misidentified, per Kovblyuk & Tuneva, 2009: 159).
T. m. Platnick & Murphy, 1984: 22, f. 51-54 (Dmf).
T. m. Chatzaki, Thaler & Mylonas, 2003: 53, f. 22-23, 28 (mf).
T. m. Kovblyuk, 2005a: 143, f. 10-14 (mf).
T. m. Kovblyuk & Tuneva, 2009: 159, f. 17-21 (mf).

mf miniglossus Levy, 2009....................Israel [urn:lsid:amnh.org:spidersp:042300]
T. m. Levy, 2009: 27, f. 59-62 (Dmf).

mf minutus Crespo, 2010....................Portugal [urn:lsid:amnh.org:spidersp:043618]
T. m. Crespo, in Crespo & Mendes, 2010: 588, f. 1-9 (Dmf).

mf mutabilis (Simon, 1878)....................Mediterranean [urn:lsid:amnh.org:spidersp:028034]
Prosthesima m. Simon, 1878a: 46, pl. 14, f. 7 (Dmf).
Zelotes m. Simon, 1914a: 153, 213, f. 282, 284 (mf).
Zelotes m. Jézéquel, 1962b: 597, f. 1 (f).
Zelotes microbarbatus Marinaro, 1967: 693, f. 10-11 (Dm).
T. m. Platnick & Murphy, 1984: 18, f. 39-42 (Tmf from Zelotes, S).

mf pedestris (C. L. Koch, 1837) *....................Europe to Azerbaijan [urn:lsid:amnh.org:spidersp:028035]
Melanophora p. C. L. Koch, 1837b: 17 (D).
Melanophora p. C. L. Koch, 1839a: 82, f. 489 (Dm).
Drassus p. Blackwall, 1861a: 442 (Df).
Drassus p. O. P.-Cambridge, 1861b: 7558.
Melanophora p. L. Koch, 1866: 178, pl. 7, f. 115-116 (f).
Melanophora p. Menge, 1872: 312, pl. 56, f. 180 (mf).
Prosthesima p. Simon, 1878a: 50.
Prosthesima p. Becker, 1896: 242, pl. 16, f. 3 (mf).
Prosthesima p. Chyzer & Kulczyn'ski, 1897: 208, pl. 7, f. 47, 49 (mf).
Prosthesima p. Bösenberg, 1902: 310, pl. 29, f. 453 (mf).
Zelotes p. Simon, 1914a: 152, 213, f. 279 (f).
Zelotes p. Reimoser, 1937a: 31, f. 56, 73 (mf).
Zelotes p. Tullgren, 1946: 118, f. 34A, pl. 19, f. 246-248 (mf).
Zelotes p. Locket & Millidge, 1951: 107, f. 56A-B, G (mf).
Zelotes p. Jézéquel, 1962a: 521, f. 1 (f).
Zelotes p. Braendegaard, 1966: 111, f. 94-95 (mf).
Zelotes p. Miller, 1967: 271, pl. III, f. 16, pl. VI, f. 6 (mf).
Zelotes p. Miller, 1971: 85, pl. VII, f. 22-23 (mf).
T. p. Platnick & Murphy, 1984: 4, f. 3-6 (Tmf from Zelotes).
Zelotes p. Grimm, 1985: 212, f. 4, 223-226 (mf).
Zelotes p. Roberts, 1985: 72, f. 26a (mf).
Zelotes p. Heimer & Nentwig, 1991: 440, f. 1158 (mf).
Zelotes p. Noordam, 1992: 9 (f).
Zelotes p. Roberts, 1995: 110, f. (mf).
T. p. Levy, 1998c: 102, f. 15-18 (mf).
Zelotes p. Roberts, 1998: 114, f. (mf).
T. p. Senglet, 2004: 99, f. 41, 43-44 (mf).
T. p. Kovblyuk, 2005a: 140, f. 1-4 (mf).
T. p. Almquist, 2006: 421, f. 363a-i (mf).
T. p. Murphy, 2007: 37, f. 190-191 (mf).

f ravidus (L. Koch, 1875)....................Ethiopia [urn:lsid:amnh.org:spidersp:028036]
Prosthesima ravida L. Koch, 1875b: 48, pl. 5, f. 4 (Df).
T. r. Platnick & Murphy, 1984: 17, f. 17-18 (Tf from Zelotes).

mf stubbsi Platnick & Murphy, 1984....................Greece, Cyprus [urn:lsid:amnh.org:spidersp:042299]
T. s. Platnick & Murphy, 1984: 9, f. 15-16 (Dm).
T. s. Levy, 1998c: 106, f. 23-24 (m).
T. adriaticus Chatzaki, Thaler & Mylonas, 2003: 54, f. 36-37 (m, misidentified).
T. s. Levy, 2009: 25, f. 57-58 (removed m from S of T. adriaticus, contra Chatzaki, Thaler & Mylonas, 2003: 54, Df).
T. s. Chatzaki, 2010b: 48, f. 6-12 (mf).